Cyperus niger |
Cyperus granitophilus |
|
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black flatsedge, brown cyperus |
granite flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, annual, cespitose, with roots fibrous. |
Culms | solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
trigonous, glabrous. |
Leaves | (2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
V-shaped to flat, (2–)4–11 cm × 3–4 mm. |
Inflorescences | spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
rays absent; bracts 2–4, ± horizontal, flat, 1–6.5 cm × 2–3 mm; rachilla persistent, wingless. |
Spikelets | (3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
10–40, compressed, ovoid to linear-lanceoloid, quadrangular, 5–9 × 2–3 mm; floral scales deciduous, 5–20, spreading, pale greenish to reddish brown, laterally (5–)7–9(–13)-ribbed, ovate, 3–4 × 1–2 mm, apex straight, excurved mucronate, or cuspidate. |
Flowers | stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
stamens 1–2; filaments 1.5 mm; anthers 0.8–1 mm, connective apex reddish yellow, less than 0.1 mm; styles 0.8–1 mm; stigmas 1–1.2 mm. |
Achenes | brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
brown to reddish brown, broadly ellipsoid to obovoid, 0.8–1.2 × 0.5–0.6 mm, base ± cuneate, apex truncate-obtuse, surfaces puncticulate. |
Cyperus niger |
Cyperus granitophilus |
|
Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Swamps, ditches, wet pastures, seeps in montane forests | Thin soil over outcrops of granite or sandstone of the Piedmont and inner coastal plain |
Elevation | 0–1200 m (0–3900 ft) | 50–400 m (200–1300 ft) |
Distribution |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
|
AL; GA; NC; SC; TN; VA
|
Discussion | Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus granitophilus is documented as an autotetraploid derivative of C. squarrosus (L. W. Garoni and W. H. Murdy 1964). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 165. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) | McVaugh: Castanea 2: 103, figs. 4–8. (1937) |
Web links |