Cyperus niger |
Cyperus fuscus |
|
---|---|---|
black flatsedge, brown cyperus |
brown flatsedge, brown galingale |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, annual, cespitose. |
Culms | solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
trigonous, 2–30 cm × 0.6–1.1 mm, glabrous. |
Leaves | (2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
flat, 4–10 cm × 2–4 mm. |
Inflorescences | spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
rays 1–3, 0.2–1.5 cm; 2d order rays present in robust plants, to 5 mm; bracts 2–3, longest erect, others spreading, 1–20 cm × 1.5–3 mm. |
Spikelets | (3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
3–12, narrowly ellipsoid, flattened, 3–7 × 0.9–1.2 mm; floral scales 8–12(–16), laterally purplish brown, medially greenish yellow, 3-ribbed medially, orbiculate, 0.9–1.1 × 1 mm, apex mucronate. |
Flowers | stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
stamen 1; anthers ellipsoid, 0.2 mm, connective not prolonged; styles 0.3–0.4 mm; stigmas 0.3 mm. |
Achenes | brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
light brown, ellipsoid, 0.7–0.9 × 0.4 mm, base barely stipelike to narrowly cuneate, apex acute, surfaces glabrous. |
Cyperus niger |
Cyperus fuscus |
|
Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Swamps, ditches, wet pastures, seeps in montane forests | Damp, disturbed soils, emergent shorelines, puddles |
Elevation | 0–1200 m (0–3900 ft) | 0–1700 m (0–5600 ft) |
Distribution |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
|
CA; CT; MA; MD; MO; NE; NJ; NV; PA; SD; VA; ON; QC; Eurasia [Introduced in North America]
|
Discussion | Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A native of temperate Eurasia, Cyperus fuscus is intermittently adventive and locally established 35°–45° N latitude. The report from New York (M. L. Fernald 1950) is based on a misidentification of C. diandrus (R. S. Mitchell and G. C. Tucker 1997). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 157. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) | Linnaeus: Sp. Pl. 1: 46. (1753) |
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