Cyperus niger |
Cyperus fugax |
|
---|---|---|
black flatsedge, brown cyperus |
withering flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, annual, cespitose. |
Culms | solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
(1–)10–30(–60) together, roundly trigonous proximally, trigonous distally, (0.5–)3–6(–12) cm × (0.2–)0.4–0.8 mm, glabrous. |
Leaves | (2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
V-shaped, (1–)3–5(–10) cm × (0.5–)1–2 mm. |
Inflorescences | spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
spikes loosely ovoid, 8–15(–25) mm wide; rachis 1–3 mm; rays (1–)2–4(–6), (0.5–)1–3(–6) cm; 2d order rays absent; bracts (1–)2–4(–5), ascending at 30–45°, V-shaped, (1–)2.5–6(–11) cm × 0.5–1.8 mm. |
Spikelets | (3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
(2–)4–6(–10), linear-lanceoloid, strongly flattened, (5–)8–12(–18) × 1.2–1.5 (1.8) mm; floral scales (5–)10–16(–32), laterally clear, stramineous or light reddish brown, medially green, laterally ribless, medially 3(–5)-ribbed, ovate, (1.2–)1.3–1.6(–2) × 1–1.2(–1.4) mm, apex obtuse, mucronate to minutely mucronulate. |
Flowers | stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
stamens 2; anthers 0.1–0.2 mm; styles 0.1–0.3(–0.4) mm; stigmas 0.3–0.4 mm. |
Achenes | brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
light to dark brown, glossy or iridescent, stipitate, oblong-obovoid, 0.9–1.1 × 0.4–0.5 mm, stipe 0.1 mm, apex truncate, apiculate, surfaces punctate. |
Cyperus niger |
Cyperus fugax |
|
Phenology | Fruiting summer. | Fruiting early summer. |
Habitat | Swamps, ditches, wet pastures, seeps in montane forests | Wet, disturbed sandy soils in full sun |
Elevation | 0–1200 m (0–3900 ft) | 0–200 m (0–700 ft) |
Distribution |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
|
LA; TX; Central America; West Indies (Cuba); South America (Ecuador) |
Discussion | Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus fugax is known from Louisiana from an early nineteeth century collection from “western Louisiana” (Hale s.n., TCD). Cyperus fugax has been treated as a form of C. polystachyos (G. Kükenthal 1935–1936; A. B. Ayers 1946); nevertheless, specific status is appropriate. The most conspicuous differences are the annual habit and small size (less than 12 cm) of C. fugax; C. polystachyos is a perennial of moderate size (mostly taller than 20 cm). Other differences are summarized below. Cyperus polystachyos is characterized as a shortly rhizomatous perennial; floral scales oblong, 1.8–2.4 mm, closely imbricate; styles 0.6–1 mm; stigmas 1.4–2 mm; anthers 0.6–0.8 mm; achenes substipitate to cuneate, slightly compressed laterally. Cyperus fugax is characterized as a nonrhizomatous annual; floral scales ovate, 1.3–1.6 mm, loosely imbricate; styles 0.1–0.2 mm; stigmas 0.3–0.4 mm; filaments 1.2–1.4 mm; anthers 0.1–0.2 mm; achenes stipitate, strongly compressed laterally. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 159. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | C. inconspicuus, C. tenellus |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) | Liebmann: Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., ser. 5, 2: 196. (1850) |
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