Cyperus niger |
Cyperus flavicomus |
|
---|---|---|
black flatsedge, brown cyperus |
white-edge flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, annual, cespitose or single-stemmed. |
Culms | solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
trigonous, easily compressed, 30–75 cm × 1–5 mm, basally soft, glabrous. |
Leaves | (2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
1–3, usually withered at anthesis, V-shaped, 5–35 cm × 2–8 mm. |
Inflorescences | spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
spikes loosely ovoid, 5–25 × 8–22 mm; rays (3–)5–11, 0.4–20 cm; 2d order rays occasionally present, 5–28 mm; bracts 3–7, horizontal to ascending at 30°, (2–)8–35 cm × (1–)4–10 mm. |
Spikelets | (3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
6–60, linear to lanceoloid, 7–30 × 2–3 mm, margins appearing serrate; floral scales 6–24, loosely overlapping, light brown to reddish brown, with conspicuous clear border, medially greenish, 5–7-ribbed, ovate-obovate, (1.4–)1.7–2.3 × 1.4–2 mm, apex acute to obtuse. |
Flowers | stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
stamens 2–3; anthers 0.4 mm, connective apex reddish, to 0.1 mm; styles 0.1–0.3 mm; stigmas 0.7–1.5 mm. |
Achenes | brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
jet black to dark reddish brown, sessile, obovoid, broadly rounded, 1.2–1.6 × 0.6–1.1 mm, surfaces minutely punctate. |
Cyperus niger |
Cyperus flavicomus |
|
Phenology | Fruiting summer. | Fruiting summer–early fall. |
Habitat | Swamps, ditches, wet pastures, seeps in montane forests | Emergent shorelines, roadsides ditches, marshes |
Elevation | 0–1200 m (0–3900 ft) | 0–200 m (0–700 ft) |
Distribution |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
|
AL; AZ; FL; GA; KY; LA; MO; MS; NC; NM; PA; SC; TX; VA; Mexico; Central America; South America; Asia; Africa [Introduced in North America]
|
Discussion | Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus flavicomus has long been called C. albomarginatus (see G. C. Tucker 1994). The closely related (perhaps not distinct) C. macrostachyos Lamarck, a pantropical subspecies, has been reported from Central America; the relationship of the two species needs further study. Cyperus flavicomus has been introduced in New York; it has not persisted there (R. S. Mitchell and G. C. Tucker 1997). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 159. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | C. albomarginatus, C. albomarginatus var. sabulosus, C. sabulosus, Pycreus albomarginatus, Pycreus sabulosus |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) | Michaux: Fl. Bor.-Amer. 1: 27. (1803) |
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