Cyperus niger |
Cyperus flavescens |
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black flatsedge, brown cyperus |
pale flatsedge, yellow flatsedge |
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Habit | Herbs, perennial, rhizomatous. | Herbs, annual, cespitose. |
Culms | solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
trigonous, (2–)4–30 cm × 0.4–2 mm, glabrous. |
Leaves | (2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
1–5, (blades often absent, base of culm with 1 reddish sheath bearing minute blade tooth 1–2 mm), (3–)10–18 cm × (0.5–)1.5–2(–2.6) mm. |
Inflorescences | spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
spikes 1–3, ovoid or ± digitate, 10–30 × 8–26 mm; rays 1–4(–6), 0.5–3(–9) cm; bracts 2–3, approximately horizontal, 1–12 cm × 0.5–2.5 mm. |
Spikelets | (3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
1–6, compressed, oblong-lanceoloid, 5–15 × (2–)2.3–2.8 mm; floral scales (4–)8–24, closely imbricate, laterally yellow to yellowish brown, margins light brown to clear, medially green, laterally ribless, medially 2–3-ribbed, 2-keeled basally, ovate, (1.5–)1.8–2.2 × 1.8 mm, apex obtuse. |
Flowers | stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
stamens 3; anthers 0.4 mm, connectives not prolonged; styles 0.5–1 mm; stigmas 0.5–0.8 mm. |
Achenes | brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
jet black to reddish brown, slightly stipitate, obovoid, 1–1.2 × 0.8 mm, apex apiculate, surface with network of rectangular longitudinally elongate cells and transverse undulations. |
Cyperus niger |
Cyperus flavescens |
|
Phenology | Fruiting summer. | Fruiting summer–early fall. |
Habitat | Swamps, ditches, wet pastures, seeps in montane forests | Damp, often disturbed soils |
Elevation | 0–1200 m (0–3900 ft) | 0–800 m (0–2600 ft) |
Distribution |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
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AL; AR; CA; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; ON; South America; West Indies; Eurasia; Africa
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Discussion | Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
North American populations have been distinguished as Cyperus flavescens var. poiformis; their taxonomic separation was not supported by M. L. Corcoran’s (1941) careful study. Plants of Mexico and Central America consistently have purplish rather than yellowish floral scales and may be recognized as C. flavescens var. piceus (Liebmann) Fernald (G. C. Tucker 1983, 1994). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 162. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycreus | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | C. flavescens var. poiformis, C. poiformis, Pycreus flavescens |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) | Linnaeus: Sp. Pl. 1: 46. (1753) |
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