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black flatsedge, brown cyperus

cyperus, flatsedge, galingale, nutsedge, umbrella-sedge

Habit Herbs, perennial, rhizomatous. Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous.
Culms

solitary or close together, trigonous, 5–40(–95) cm, glabrous.

solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles.

Leaves

(2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm.

usually basal;

ligules absent;

blades keeled abaxially, flat, V-, or inversely W-shaped in cross section.

Inflorescences

spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.;

rachis 1–2 mm, or absent;

rays 1–2, 3–45 mm;

if rays absent, inflorescence sessile, dense, 6–20 mm diam;

bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm.

terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet;

spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts;

involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike.

Spikelets

(3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm;

floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse.

scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous).

Flowers

stamens 2;

anthers 0.6–0.8 mm, connectives not prolonged;

styles 0.4–0.6 mm;

stigmas 0.7–2.1 mm.

bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base deciduous or persistent;

stigmas 2–3.

Achenes

brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate.

biconvex, flattened, or trigonous.

Cyperus niger

Cyperus

Phenology Fruiting summer.
Habitat Swamps, ditches, wet pastures, seeps in montane forests
Elevation 0–1200 m (0–3900 ft)
Distribution
from FNA
AZ; CA; NM; OK; TX; Mexico; Central America; South America
[WildflowerSearch map]
[BONAP county map]
from USDA
Pantemperate and tropical
[BONAP county map]
Discussion

Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 600 (96 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas 2 (3 in C. serotinus); achenes biconvex.
→ 2
1. Stigmas 3; achenes trigonous, plano-convex, or terete.
→ 3
2. Achenes laterally flattened, borne with edge toward rachilla.
subg. Pycreus
2. Achenes dorsiventrally flattened, borne with face toward rachilla.
subg. Juncellus
3. Spikelets borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads.
subg. Pycnostachys
3. Spikelets borne in spikes on conspicuous rachis.
→ 4
4. Rachilla continuous or articulate only at base.
subg. Cyperus
4. Rachilla articulate at base of each scale, mature spikelet disarticulating into segments consisting of scale, internode, and rachilla wings.
subg. Diclidium
Source FNA vol. 23. FNA vol. 23, p. 141. Authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.
Parent taxa Cyperaceae > Cyperus > subg. Pycreus Cyperaceae
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Subordinate taxa
C. subg. Cyperus, C. subg. Diclidium, C. subg. Juncellus, C. subg. Pycnostachys, C. subg. Pycreus
Synonyms C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus
Name authority Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754)
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