Cyperus laevigatus |
Cyperus virens |
|
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smooth cyperus, smooth flatsedge, smooth nutgrass |
green flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. |
Culms | single or close together, trigonous to ± terete, (1–)8–30(–60) cm × (0.3–)0.6–1.5(–2.2) mm, glabrous. |
trigonous, (20–)60–70(–100) cm × 2.4–6.9 mm, scabrid on angles. |
Leaves | blades present or reduced to sheaths, base of culm covered with reddish sheath 5–25 mm, distally short toothed 1–2 mm, or leaf blades involute, 1–7 cm × 0.3–1(–1.6) mm. |
4–12, flat to V-shaped, with readily visible cross ribs especially on abaxial surface, 20–50 cm × 3–14 mm. |
Inflorescences | heads appearing lateral, spikes ± digitate, 5–15 × 5–15 mm; rays absent, sometimes 1, to 2 mm; bracts (1–)2(–3), longer bracts erect, appearing as continuation of culm, shorter bract(s) horizontal or ascending, longer bracts (1–)2–8(–12) cm × 0.5–3 mm, shorter (0.3–)1–4.5 cm × 0.2–1 mm. |
heads digitate to hemispheric, 1–3.5 cm diam.; rays 6–12, 1–9(–14) cm; 2d order rays often present, (0.5–)1–3 cm; 3d order rays occasionally present, 5–12 mm; bracts 4–8, horizontal to ascending at 30°, with prominent cross ribs,V-shaped, (1.5–)3–50(–75) cm × 0.5–13 mm; 2d order bracts 3–10 × 1–2 mm. |
Spikelets | (1–)2–7(–14), whitish green to reddish, compressed-turgid, ellipsoid to oblong-lanceoloid, 4–7(–12) × (1.6–)2–3(–3.6) mm; floral scales (8–)12–22, off-white, whitish with red speckles, or with single red spot 0.5 mm wide, medially whitish, oblong-laterally ribless or weakly 2-ribbed, medially weakly 1–3-ribbed, obovate to orbiculate, 1.5–2 × (1.3–)1.6–2 mm. |
10–40(–50), oblong to linear-lanceoloid, 5–18 × 2–3.2 mm; floral scales 10–40, pale grayish brown, or greenish, proximally greenish or brownish, oblanceolate, 2-keeled, weakly to distinctly 1-ribbed, proximal half 2-ribbed, (1.3–)1.5–2(–2.4) × 0.9–1.4 mm, apex acute (sometimes mucronulate), apically glabrous, occasionally scabridulous. |
Flowers | anthers (0.5–)0.7–1 mm, connective apex reddish, 0.1–0.2 mm; styles 0.8–1.2 mm; stigmas 0.7–1 mm. |
stamens 1–2; anthers 0.8–1.4 mm; styles 0.8–1 mm; stigmas 0.6–0.8 mm. |
Achenes | gray, glossy, oblong-ellipsoid to ovoid, (1.2–)1.3–1.5(–1.8) × (0.7–)0.8–1(–1.2) mm, base cuneate to stipelike, apex apiculate, surfaces finely reticulate to papillate. |
brown, slightly to distinctly stipitate, ellipsoid (2.7–4.1 times as long as wide), (0.9–)1.2–1.5 × 0.3–0.4 mm, apical beak 0.1–0.5 mm, surfaces glabrous. |
Cyperus laevigatus |
Cyperus virens |
|
Phenology | Fruiting summer. | Fruiting spring–fall. |
Habitat | Alkaline grasslands, hotsprings, ditches, brackish marshes | Wet pastures, marshes, roadside ditches |
Elevation | 0–1000 m (0–3300 ft) | |
Distribution |
AZ; CA; NC; TX; Mexico; South America; West Indies; Europe; Asia; Africa
|
AL; FL; GA; LA; MS; NC; SC; TX; Mexico; Central America; South America
|
Discussion | Cyperus laevigatus is one of only two species of subg. Juncellus in the New World. It can be distinguished from all other New World species of Cyperus by its two stigmas and disclike, strongly flattened achene, which is borne with its face toward the rachilla. Species of subg. Pycreus, especially C. niger, have been misidentified as C. laevigatus, probably because of the dark, shiny spikelets. All species of subg. Pycreus have biconvex achenes like C. laevigatus; in species of subg. Pycreus the achene is attached with an edge toward the rachilla; in C. laevigatus the achene faces the rachilla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus virens was found as waif in California in the 1800s (M. F. Denton 1978b; G. C. Tucker 1993b). Cyperus virens is readily distinguished from other species of subg. Pycnostachys by its trigonous culms, scabrid angles, and leaves and inflorescence bracts conspicuously septate by numerous cross ribs between the main ribs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 164. | FNA vol. 23, p. 153. |
Parent taxa | Cyperaceae > Cyperus > subg. Juncellus | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | Acorellus laevigatus, Chlorocyperus laevigatus, Juncellus laevigatus, Pycreus laevigatus | |
Name authority | Linnaeus: Mant. Pl. 2: 179. (1771) | Michaux: Fl. Bor.-Amer. 1: 28. (1803) |
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