FNA: Cyperus involucratus vs. Cyperus sphaerolepis

	Cyperus involucratus	Cyperus sphaerolepis
	umbrella-plant	Rusby's flatsedge, Rusby's sedge
Habit	Herbs, perennial, cespitose, rhizomatous.	Herbs, perennial, rhizomatous.
Culms	trigonous, 30–150 cm × 1–5(–8) mm.	basally cormlike, trigonous, (8–)15–40(–60) cm × (0.4–)0.8–1.5(–2) mm, glabrous or sparsely to densely scabridulous to minutely scabrid on angles or usually on surface, just proximal to bracts or over distal few centimeters.
Leaves	bladeless.	(1–)3–4, V-shaped, (7–)15–25(–40) cm × 1–3(–5) mm, sparsely minutely scabridulous on margins and keel in distal 1/3, otherwise glabrous.
Inflorescences	heads digitate, 15–30(–36) mm diam.; rays (14–)20–22, (2–)5–12(–20) cm; 2d order rays 0.3–3(–4) cm; 3d order rays sometimes present, 0.3–2.5 cm; bracts (4–)18–22, ± horizontal, flat, 15–27 cm × (1.5–)8–12 mm.	spikes 1–3, sessile (occasionally 1 spike on peduncle 5–20 cm, produced from axil of distalmost leaf), loosely cylindric to oblong-ovoid, (9–)15–30(–45) × (3–)8–15(–18) mm; bracts 2–3(–7), longest erect, or very nearly so, others spreading, (1–)3–20(–26) cm × (0.5–)1–3(–5) mm; rays (1–)2–6(–11), (1–)2–7(–12) cm, in some depauperate plants rays less than 1 cm; rachilla ± deciduous, wingless.
Spikelets	8–20, ovoid to linear-lanceoloid, compressed, 5–25 × 1.5–2 mm; floral scales 8–28, laterally whitish or light brown, ± hyaline, medially light brown, laterally ribless, medially 3-ribbed, 2-keeled in proximal 30–60%, deltate-ovate, 1.6–2.4 × (1–)1.2–1.5(–1.7) mm, apex acute.	(4–)8–25(–35), oblong-ellipsoid to linear, compressed-quadrangular, (2–)7–11(–15) × (2–)2.4–3.2(–3.5) mm; floral scales (1–)4–8(–14), laterally dull whitish, stramineous, or light brown, or reddish brown to deep red, often red-dotted especially near keel, medially greenish, broadly ovate-deltate to ± orbiculate, laterally strongly 3(–4)-ribbed, medially weakly 1–3-ribbed, (1.6–)1.8–2.4 × (1.8–)2.2–2.8 mm, margins clear, erose especially apically, apex obtuse, tipped with straight mucro 0.1–0.2(–0.3) mm.
Flowers	stamens 3; anthers 0.7–1 mm; styles 0.5–1 mm; stigmas 0.6–1 mm.	anthers 0.4–0.6 mm; styles 0.4(–0.8) mm; stigmas 0.6–1 mm.
Achenes	brown, sessile or stipitate, broadly ellipsoid, 0.6–0.8 × 0.4–0.6 mm, stipe if present to 0.1 mm, apex obtuse, apiculate, surfaces puncticulate.	finely reticulate with isodiametric cells, this layer gradually wearing away to reveal inner brown to reddish brown layer, stipitate, ovoid to obovoid, 1.4–1.6(–1.9) × 1–1.3(–1.4) mm, base cuneate to ± attenuate, stipe 0.1–0.2 × 0.2 mm, apex obtuse, slightly apiculate, inner layer papillose.

	Cyperus involucratus	Cyperus sphaerolepis
Phenology	Fruiting early summer–fall.	Fruiting summer.
Habitat	Damp, disturbed soils, ditches, stream banks	Clearings in montane forests
Elevation	0–100(–800) m [0–300(–2600) ft]	1000–2500 m [3300–8200 ft]
Distribution	CA; FL; LA; TX; e Africa [Introduced in North America] [BONAP county map]	AZ; NM; TX; Mexico [BONAP county map]
Discussion	Cyperus involucratus has been collected in New York (R. S. Mitchell and G. C. Tucker 1997). Cyperus involucratus is widely cultivated as a water plant in greenhouses and outdoors in warm-temperate or tropical climates. It has long been misidentified in the flora as C. alternifolius Linnaeus, an endemic of Madagascar (G. C. Tucker 1983). (Discussion copyrighted by Flora of North America; reprinted with permission.)	Cyperus sphaerolepis has long been known by the synonym C. rusbyi and has been treated as a variety of C. fendlerianus. Specific status for C. sphaerolepis is supported by B. G. Marcks (1972) and G. C. Tucker (1994). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23, p. 150. Treatment authors: Gordon C. Tucker, Brian G. Marcks, J. Richard Carter *.	FNA vol. 23, p. 177. Treatment authors: Gordon C. Tucker, Brian G. Marcks, J. Richard Carter *.
Parent taxa	Cyperaceae > Cyperus > subg. Pycnostachys	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms		C. fendlerianus var. debilis, C. fendlerianus var. leucolepis, C. leucolepis, C. rusbyi, C. schweinitzii var. debilis
Name authority	Rottbøll: Descr. Pl. Rar., 22. (1772)	Boeckeler: Linnaea 35: 609. (1868)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos, LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus sphaerolepis

umbrella-plant

Rusby's flatsedge, Rusby's sedge

Habit

Herbs, perennial, cespitose, rhizomatous.

Herbs, perennial, rhizomatous.

Culms

trigonous, 30–150 cm × 1–5(–8) mm.

basally cormlike, trigonous, (8–)15–40(–60) cm × (0.4–)0.8–1.5(–2) mm, glabrous or sparsely to densely scabridulous to minutely scabrid on angles or usually on surface, just proximal to bracts or over distal few centimeters.

Leaves

bladeless.

(1–)3–4, V-shaped, (7–)15–25(–40) cm × 1–3(–5) mm, sparsely minutely scabridulous on margins and keel in distal 1/3, otherwise glabrous.

Inflorescences

heads digitate, 15–30(–36) mm diam.;

rays (14–)20–22, (2–)5–12(–20) cm; 2d order rays 0.3–3(–4) cm; 3d order rays sometimes present, 0.3–2.5 cm;

bracts (4–)18–22, ± horizontal, flat, 15–27 cm × (1.5–)8–12 mm.

spikes 1–3, sessile (occasionally 1 spike on peduncle 5–20 cm, produced from axil of distalmost leaf), loosely cylindric to oblong-ovoid, (9–)15–30(–45) × (3–)8–15(–18) mm;

bracts 2–3(–7), longest erect, or very nearly so, others spreading, (1–)3–20(–26) cm × (0.5–)1–3(–5) mm;

rays (1–)2–6(–11), (1–)2–7(–12) cm, in some depauperate plants rays less than 1 cm;

rachilla ± deciduous, wingless.

Spikelets

8–20, ovoid to linear-lanceoloid, compressed, 5–25 × 1.5–2 mm;

floral scales 8–28, laterally whitish or light brown, ± hyaline, medially light brown, laterally ribless, medially 3-ribbed, 2-keeled in proximal 30–60%, deltate-ovate, 1.6–2.4 × (1–)1.2–1.5(–1.7) mm, apex acute.

(4–)8–25(–35), oblong-ellipsoid to linear, compressed-quadrangular, (2–)7–11(–15) × (2–)2.4–3.2(–3.5) mm;

floral scales (1–)4–8(–14), laterally dull whitish, stramineous, or light brown, or reddish brown to deep red, often red-dotted especially near keel, medially greenish, broadly ovate-deltate to ± orbiculate, laterally strongly 3(–4)-ribbed, medially weakly 1–3-ribbed, (1.6–)1.8–2.4 × (1.8–)2.2–2.8 mm, margins clear, erose especially apically, apex obtuse, tipped with straight mucro 0.1–0.2(–0.3) mm.

Flowers

stamens 3;

anthers 0.7–1 mm;

styles 0.5–1 mm;

stigmas 0.6–1 mm.

anthers 0.4–0.6 mm;

styles 0.4(–0.8) mm;

stigmas 0.6–1 mm.

Achenes

brown, sessile or stipitate, broadly ellipsoid, 0.6–0.8 × 0.4–0.6 mm, stipe if present to 0.1 mm, apex obtuse, apiculate, surfaces puncticulate.

finely reticulate with isodiametric cells, this layer gradually wearing away to reveal inner brown to reddish brown layer, stipitate, ovoid to obovoid, 1.4–1.6(–1.9) × 1–1.3(–1.4) mm, base cuneate to ± attenuate, stipe 0.1–0.2 × 0.2 mm, apex obtuse, slightly apiculate, inner layer papillose.

Cyperus involucratus

Cyperus sphaerolepis

Phenology

Fruiting early summer–fall.

Fruiting summer.

Habitat

Damp, disturbed soils, ditches, stream banks

Clearings in montane forests

Elevation

0–100(–800) m [0–300(–2600) ft]

1000–2500 m [3300–8200 ft]

Distribution

CA; FL; LA; TX; e Africa [Introduced in North America]

[BONAP county map]

AZ; NM; TX; Mexico

[BONAP county map]

Discussion

Cyperus involucratus has been collected in New York (R. S. Mitchell and G. C. Tucker 1997).

Cyperus involucratus is widely cultivated as a water plant in greenhouses and outdoors in warm-temperate or tropical climates. It has long been misidentified in the flora as C. alternifolius Linnaeus, an endemic of Madagascar (G. C. Tucker 1983).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus sphaerolepis has long been known by the synonym C. rusbyi and has been treated as a variety of C. fendlerianus. Specific status for C. sphaerolepis is supported by B. G. Marcks (1972) and G. C. Tucker (1994).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23, p. 150. Treatment authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.

FNA vol. 23, p. 177. Treatment authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.

Parent taxa

Cyperaceae > Cyperus > subg. Pycnostachys

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

C. fendlerianus var. debilis, C. fendlerianus var. leucolepis, C. leucolepis, C. rusbyi, C. schweinitzii var. debilis

Name authority

Rottbøll: Descr. Pl. Rar., 22. (1772)

Boeckeler: Linnaea 35: 609. (1868)

Web links

Local floras: CA
Local Web sites: CalFlora, CalPhotos, LA Plants
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Flora of North America species comparison

Cyperus involucratus

Cyperus sphaerolepis

Cyperus involucratus

Cyperus sphaerolepis