Cyperus involucratus |
Cyperus papyrus |
|
---|---|---|
umbrella-plant |
Egyptian paper-reed, papyrus |
|
Habit | Herbs, perennial, cespitose, rhizomatous. | Herbs, perennial, rhizomatous, stout. |
Culms | trigonous, 30–150 cm × 1–5(–8) mm. |
roundly trigonous, 300–500 cm × 15–45 mm, glabrous. |
Leaves | bladeless. |
bladeless. |
Inflorescences | heads digitate, 15–30(–36) mm diam.; rays (14–)20–22, (2–)5–12(–20) cm; 2d order rays 0.3–3(–4) cm; 3d order rays sometimes present, 0.3–2.5 cm; bracts (4–)18–22, ± horizontal, flat, 15–27 cm × (1.5–)8–12 mm. |
spikes loosely cylindric, 10–20 × 6–10 mm; rays 40–100, drooping or arching, slender, 10–30 cm; 2d order rays 8–20 cm; bracts 4–10, ± erect, V-shaped, 3–8 cm × 4–15 mm; 2d order bracts 2–5, (1.5–)4–16 cm × 0.5–2 mm; rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide. |
Spikelets | 8–20, ovoid to linear-lanceoloid, compressed, 5–25 × 1.5–2 mm; floral scales 8–28, laterally whitish or light brown, ± hyaline, medially light brown, laterally ribless, medially 3-ribbed, 2-keeled in proximal 30–60%, deltate-ovate, 1.6–2.4 × (1–)1.2–1.5(–1.7) mm, apex acute. |
6–30, slightly compressed, linear, ± quadrangular, 6–10 × 0.8–1 mm; floral scales 6–16, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate-elliptic, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse. |
Flowers | stamens 3; anthers 0.7–1 mm; styles 0.5–1 mm; stigmas 0.6–1 mm. |
anthers 0.8–1 mm (connectives prolonged beyond anther as red subulate appendage 0.2–0.5 mm, apex usually setose); styles 0.2–0.4 mm; stigmas 0.8–1.2 mm. |
Achenes | brown, sessile or stipitate, broadly ellipsoid, 0.6–0.8 × 0.4–0.6 mm, stipe if present to 0.1 mm, apex obtuse, apiculate, surfaces puncticulate. |
pale brown, sessile, oblong, 0.8–1 × 0.4 mm, apex scarcely apiculate, surfaces puncticulate. |
Cyperus involucratus |
Cyperus papyrus |
|
Phenology | Fruiting early summer–fall. | Fruiting summer. |
Habitat | Damp, disturbed soils, ditches, stream banks | Stream banks, marshes |
Elevation | 0–100(–800) m [0–300(–2600) ft] | 0–30 m [0–100 ft] |
Distribution |
CA; FL; LA; TX; e Africa [Introduced in North America] |
FL; s Europe; sw Asia; Africa [Introduced in North America] |
Discussion | Cyperus involucratus has been collected in New York (R. S. Mitchell and G. C. Tucker 1997). Cyperus involucratus is widely cultivated as a water plant in greenhouses and outdoors in warm-temperate or tropical climates. It has long been misidentified in the flora as C. alternifolius Linnaeus, an endemic of Madagascar (G. C. Tucker 1983). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus papyrus is conspicuous in the field by its great height, leafless culm, and open, hemispheric inflorescence with drooping rays. Ancient Egyptians, Greeks, and Romans prepared paper from the pith of the culms of Cyperus papyrus, which is abundant along the Nile River. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 150. | FNA vol. 23, p. 173. |
Parent taxa | ||
Sibling taxa | ||
Name authority | Rottbøll: Descr. Pl. Rar., 22. (1772) | Linnaeus: Sp. Pl. 1: 47. (1753) |
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