Cyperus involucratus |
Cyperus odoratus |
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umbrella flatsedge, umbrella-plant |
fragrant flatsedge, rusty flat sedge, scented sedge |
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Habit | Herbs, perennial, cespitose, rhizomatous. | Herbs, annual (or short-lived perennial). |
Culms | trigonous, 30–150 cm × 1–5(–8) mm. |
trigonous, (4–)10–50(–130) cm × (0.5–)1–4 mm. |
Leaves | bladeless. |
flanged V- or inversely W-shaped, 5–30(–60) cm × 4–12 mm. |
Inflorescences | heads digitate, 15–30(–36) mm diam.; rays (14–)20–22, (2–)5–12(–20) cm; 2d order rays 0.3–3(–4) cm; 3d order rays sometimes present, 0.3–2.5 cm; bracts (4–)18–22, ± horizontal, flat, 15–27 cm × (1.5–)8–12 mm. |
spikes 1–5(–12), loosely to densely cylindric, ovoid, or ± pyramidal, 10–20(–40) × (8–)10–30(–35) mm; rays (0–2)6–9(–12), 2–8(–13) cm; if rays absent, infloresence a single dense, capitate cluster of closely imbricate spikes; bracts (4–)5–8(–10), horizontal to ascending at 30–60°, (3–)10–25(–55) cm × 1–14 mm; rachilla hyaline or thickened bronze, carmine, or yellowish, wings clasping achene. |
Spikelets | 8–20, ovoid to linear-lanceoloid, compressed, 5–25 × 1.5–2 mm; floral scales 8–28, laterally whitish or light brown, ± hyaline, medially light brown, laterally ribless, medially 3-ribbed, 2-keeled in proximal 30–60%, deltate-ovate, 1.6–2.4 × (1–)1.2–1.5(–1.7) mm, apex acute. |
(10–)20–60(–100), linear-oblong to narrowly linear, cylindric to slightly flattened (when scale tips spreading), (5–)8–15(–38) × 0.8–1.3(–1.9) mm; floral scales (4–)8–12–(30), medially green, laterally reddish to stramineous to shiny brown or beige, medially 2–5-ribbed, laterally 1–3-ribbed, ovate to elliptic, (2–)2.2–2.8(–3.2) × (1.2–)1.4–1.6(–1.8) mm, apex entire or emarginate with mucro to 0.3 mm. |
Flowers | stamens 3; anthers 0.7–1 mm; styles 0.5–1 mm; stigmas 0.6–1 mm. |
anthers (0.2–)0.3–0.4(–0.7) mm; styles 0.4–0.7(–1) mm; stigmas (1–)1.5–3 mm. |
Achenes | brown, sessile or stipitate, broadly ellipsoid, 0.6–0.8 × 0.4–0.6 mm, stipe if present to 0.1 mm, apex obtuse, apiculate, surfaces puncticulate. |
brown, reddish brown, or black, stipitate, narrowly ellipsoid to oblong (rarely obovoid-oblong), (1–)1.2–1.5(–1.9) × 0.5–0.6(–0.75) mm, stipe 0.1–0.2 mm, apex acute to barely obtuse, surfaces finely papillose. |
Cyperus involucratus |
Cyperus odoratus |
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Phenology | Fruiting early summer–fall. | Fruiting summer–early fall. |
Habitat | Damp, disturbed soils, ditches, stream banks | Emergent shorelines, disturbed, muddy places, fresh or slightly brackish marshes |
Elevation | 0–100(–800) m (0–300(–2600) ft) | 0–1500 m (0–4900 ft) |
Distribution |
CA; FL; LA; TX; e Africa [Introduced in North America]
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AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; HI; ON; QC; Central America; South America; Asia; Africa; Australia
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Discussion | Cyperus involucratus has been collected in New York (R. S. Mitchell and G. C. Tucker 1997). Cyperus involucratus is widely cultivated as a water plant in greenhouses and outdoors in warm-temperate or tropical climates. It has long been misidentified in the flora as C. alternifolius Linnaeus, an endemic of Madagascar (G. C. Tucker 1983). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As circumscribed here, Cyperus odoratus is easily identified by its cylindric to subcylindric spikelets in which the corky rachilla of the mature spikelet disarticulates at the base of each scale. The mature spikelet breaks into segments each consisting of a scale and an internode of the rachilla clasping the achene with its corky wings. Cyperus odoratus is an exceedingly variable pantropical and warm-temperate species. Numerous segregates have been named, some of which may deserve recognition when the species is studied in detail worldwide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 150. | FNA vol. 23, p. 191. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Diclidium |
Sibling taxa | ||
Synonyms | C. acicularis, C. californicus, C. eggersii, C. engelmannii, C. ferax, C. ferax subsp. engelmannii, C. ferax subsp. speciosus, C. ferruginescens, C. haenkei, C. huarmensis, C. macrocephalus, C. michauxianus, C. odoratus var. engelmannii, C. oxycarioides, C. speciosus, C. squarrosus var. parvus, Diclidium aciculare, Diclidium odoratum, Mariscus ferax, Mariscus huarmensis, Torulinium eggersii, Torulinium ferax, Torulinium macrocephalum, Torulinium michauxianum, Torulinium odoratum | |
Name authority | Rottbøll: Descr. Pl. Rar., 22. (1772) | Linnaeus: Sp. Pl. 1: 46. (1753) |
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