Cyperus involucratus |
Cyperus niger |
|
---|---|---|
umbrella-plant |
black flatsedge, brown cyperus |
|
Habit | Herbs, perennial, cespitose, rhizomatous. | Herbs, perennial, rhizomatous. |
Culms | trigonous, 30–150 cm × 1–5(–8) mm. |
solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
Leaves | bladeless. |
(2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
Inflorescences | heads digitate, 15–30(–36) mm diam.; rays (14–)20–22, (2–)5–12(–20) cm; 2d order rays 0.3–3(–4) cm; 3d order rays sometimes present, 0.3–2.5 cm; bracts (4–)18–22, ± horizontal, flat, 15–27 cm × (1.5–)8–12 mm. |
spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
Spikelets | 8–20, ovoid to linear-lanceoloid, compressed, 5–25 × 1.5–2 mm; floral scales 8–28, laterally whitish or light brown, ± hyaline, medially light brown, laterally ribless, medially 3-ribbed, 2-keeled in proximal 30–60%, deltate-ovate, 1.6–2.4 × (1–)1.2–1.5(–1.7) mm, apex acute. |
(3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
Flowers | stamens 3; anthers 0.7–1 mm; styles 0.5–1 mm; stigmas 0.6–1 mm. |
stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
Achenes | brown, sessile or stipitate, broadly ellipsoid, 0.6–0.8 × 0.4–0.6 mm, stipe if present to 0.1 mm, apex obtuse, apiculate, surfaces puncticulate. |
brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
Cyperus involucratus |
Cyperus niger |
|
Phenology | Fruiting early summer–fall. | Fruiting summer. |
Habitat | Damp, disturbed soils, ditches, stream banks | Swamps, ditches, wet pastures, seeps in montane forests |
Elevation | 0–100(–800) m [0–300(–2600) ft] | 0–1200 m [0–3900 ft] |
Distribution |
CA; FL; LA; TX; e Africa [Introduced in North America] |
AZ; CA; NM; OK; TX; Mexico; Central America; South America |
Discussion | Cyperus involucratus has been collected in New York (R. S. Mitchell and G. C. Tucker 1997). Cyperus involucratus is widely cultivated as a water plant in greenhouses and outdoors in warm-temperate or tropical climates. It has long been misidentified in the flora as C. alternifolius Linnaeus, an endemic of Madagascar (G. C. Tucker 1983). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 150. | FNA vol. 23. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus | |
Name authority | Rottbøll: Descr. Pl. Rar., 22. (1772) | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) |
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