Cyperus involucratus |
Cyperus esculentus |
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umbrella flatsedge, umbrella-plant |
nut flatsedge, nut grass, yellow nut sedge, yellow nut-grass |
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Habit | Herbs, perennial, cespitose, rhizomatous. | Herbs, perennial, stoloniferous; stolons soft, spongy, flexible when dried, 2–8(–15) cm, bearing tubers (3–)6–11 mm diam. | ||||||||
Culms | trigonous, 30–150 cm × 1–5(–8) mm. |
trigonous, 15–60 (–100) cm × 0.6–3.4 mm, glabrous. |
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Leaves | bladeless. |
3–7, flat to V-shaped or flanged V-shaped, (6–)20–40(–80) cm × 2–4 (–6.5) mm. |
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Inflorescences | heads digitate, 15–30(–36) mm diam.; rays (14–)20–22, (2–)5–12(–20) cm; 2d order rays 0.3–3(–4) cm; 3d order rays sometimes present, 0.3–2.5 cm; bracts (4–)18–22, ± horizontal, flat, 15–27 cm × (1.5–)8–12 mm. |
spikes broadly ellipsoid to ovoid or hemispheric, (12–)18–30 × (12–)18–35 mm; rachis 4–17 mm; rays 4–10, (0.3–)2–12 cm; bracts (3–)4–5(–7), ± horizontal to ascending at 45(–75)°, V-shaped to flanged V-shaped, (1.5–)5–30 cm × 0.5–4 mm; rachilla persistent, wings hyaline, 0.3–0.5 mm wide. |
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Spikelets | 8–20, ovoid to linear-lanceoloid, compressed, 5–25 × 1.5–2 mm; floral scales 8–28, laterally whitish or light brown, ± hyaline, medially light brown, laterally ribless, medially 3-ribbed, 2-keeled in proximal 30–60%, deltate-ovate, 1.6–2.4 × (1–)1.2–1.5(–1.7) mm, apex acute. |
(3–)10–20(–28), divaricate or ascending, yellowish brown to dark brown, linear to linear-lanceoloid, compressed-quadrangular, (5–)10–20(–55) × (1.2–)1.4–2(–3) mm; floral scales persistent, 6–34, spreading or appressed, ovate-lanceolate, laterally 7–9-ribbed, laterally yellowish brown to dark brown medially brownish, reddish, or greenish, ovate, or ovate-deltate, medially 3-ribbed, 1.8–2.7(–3.4) × (1–)1.5–1.8(–2.4) mm, apex acute or subacute. |
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Flowers | stamens 3; anthers 0.7–1 mm; styles 0.5–1 mm; stigmas 0.6–1 mm. |
anthers (1–)1.2–1.5(–2.1) mm; styles (0.7–)1–1.2(–2.2) mm; stigmas (1.2–)1.8–2.3(–4) mm. |
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Achenes | brown, sessile or stipitate, broadly ellipsoid, 0.6–0.8 × 0.4–0.6 mm, stipe if present to 0.1 mm, apex obtuse, apiculate, surfaces puncticulate. |
(seldom maturing) brown, sessile, ellipsoid, (1.1–)1.3–1.5(–1.6) × 0.3–0.6(–0.8) mm, apex obtuse, surfaces puncticulate. |
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Cyperus involucratus |
Cyperus esculentus |
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Phenology | Fruiting early summer–fall. | |||||||||
Habitat | Damp, disturbed soils, ditches, stream banks | |||||||||
Elevation | 0–100(–800) m (0–300(–2600) ft) | |||||||||
Distribution |
CA; FL; LA; TX; e Africa [Introduced in North America]
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AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; BC; NB; ON; QC; nearly worldwide
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Discussion | Cyperus involucratus has been collected in New York (R. S. Mitchell and G. C. Tucker 1997). Cyperus involucratus is widely cultivated as a water plant in greenhouses and outdoors in warm-temperate or tropical climates. It has long been misidentified in the flora as C. alternifolius Linnaeus, an endemic of Madagascar (G. C. Tucker 1983). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (3 in the flora). Cyperus esculentus is a widespread and polymorphic species. Although seven varieties have been recognized (G. Kükenthal (1935–1936), recent studies based primarily on spikelet features provided support for four varieties (P. Schippers et al. 1995). Cyperus esculentus var. esculentus is restricted to the Old World. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 150. | FNA vol. 23. | ||||||||
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Rottbøll: Descr. Pl. Rar., 22. (1772) | Linnaeus: Sp. Pl. 1: 45. (1753) | ||||||||
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