Cyperus hermaphroditus |
Cyperus squarrosus |
|
---|---|---|
hermaphrodite flatsedge |
awn cyperus, awn flatsedge, bearded flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, annual, cespitose, with fibrous roots. |
Culms | trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous. |
1–20, trigonous, 2–16 cm × 0.5–0.8 mm, glabrous. |
Leaves | V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous. |
1–3, flat to V-shaped, (1–)5–10(–15) × 0.5–2.5 mm. |
Inflorescences | spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide. |
spike 1, loosely to densely ovoid to oblong, 6–20(–40) × 9–15(–20) mm; rays absent or 1–3(–6), 4–40 mm; bracts (1–)2–4, longest ± erect, V-shaped, 1–15 cm × 0.5–3 mm; rachilla ± deciduous, wingless. |
Spikelets | (10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse. |
(2–)6–20(–40), greenish to reddish brown, somewhat compressed, ovoid-lanceoloid to oblong, 2.5–10(–20) × 1.3–2.2 mm (excluding awns); floral scales deciduous, (4–)10–20(–34), greenish to stramineous or brownish red laterally, greenish medially, (5–)7–9(–11)-ribbed nearly to margins, oblong-lanceolate, (1.2–)1.3–1.8(–2.2) × (0.5–)0.7–0.8(–1) mm, apex cuspidate, excurved awn additional 0.5–1(–1.3) mm. |
Flowers | anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm. |
stamen 1, filaments 1.5 mm; anthers oblong, 0.2–0.3 mm, connective apex reddish, minute; styles 0.3–0.5 mm; stigmas 0.4–0.7 mm. |
Achenes | brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate. |
light brown to nearly black, stipitate, obovoid (occasionally linear-spatulate or linear oblong, infrequently constricted in middle), 0.7–0.8(–1.1) × (0.2–)0.3–0.4(–0.5) mm, stipe 0.05–0.1 × 0.1 mm, apex truncate, apiculate, surfaces papillose. |
Cyperus hermaphroditus |
Cyperus squarrosus |
|
Phenology | Fruiting summer–fall. | Fruiting summer. |
Habitat | Arroyos | Moist, disturbed soils, gravelly roadsides, flood plains, edges of puddles, muddy places |
Elevation | 500–1200 m (1600–3900 ft) | 0–2500 m (0–8200 ft) |
Distribution |
AZ; TX; Mexico; South America; West Indies |
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; ON; QC; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
|
Discussion | An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus squarrosus can be recognized by its small size and annual habit combined with its oblong-lanceolate floral scales bearing five to eleven conspicuous ribs and excurved awns. Some collections have been misidentified as C. acuminatus, an annual species of subg. Pycnostachys that has ovate-lanceolate, three-ribbed floral scales and digitately clustered spikelets. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 186. | FNA vol. 23, p. 165. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii | Chlorocyperus inflexus, C. aristatus, C. aristatus var. inflexus, C. aristatus var. runyonii, C. inflexus, Dichostylis aristata, Mariscus squarrosus |
Name authority | (Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916) | Linnaeus: Cent. Pl. II, 6. (1756) |
Web links |
|