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hermaphrodite flatsedge

fragrant flatsedge, rusty flat sedge, scented sedge

Habit Herbs, perennial, rhizomatous. Herbs, annual (or short-lived perennial).
Culms

trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous.

trigonous, (4–)10–50(–130) cm × (0.5–)1–4 mm.

Leaves

V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous.

flanged V- or inversely W-shaped, 5–30(–60) cm × 4–12 mm.

Inflorescences

spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis;

rays (3–)5–10(–17), (1–)5–12(–24) cm;

bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm;

rachilla deciduous, wings 0.4–0.5 mm wide.

spikes 1–5(–12), loosely to densely cylindric, ovoid, or ± pyramidal, 10–20(–40) × (8–)10–30(–35) mm;

rays (0–2)6–9(–12), 2–8(–13) cm;

if rays absent, infloresence a single dense, capitate cluster of closely imbricate spikes;

bracts (4–)5–8(–10), horizontal to ascending at 30–60°, (3–)10–25(–55) cm × 1–14 mm;

rachilla hyaline or thickened bronze, carmine, or yellowish, wings clasping achene.

Spikelets

(10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm;

distal spikelet spreading or ascending;

floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse.

(10–)20–60(–100), linear-oblong to narrowly linear, cylindric to slightly flattened (when scale tips spreading), (5–)8–15(–38) × 0.8–1.3(–1.9) mm;

floral scales (4–)8–12–(30), medially green, laterally reddish to stramineous to shiny brown or beige, medially 2–5-ribbed, laterally 1–3-ribbed, ovate to elliptic, (2–)2.2–2.8(–3.2) × (1.2–)1.4–1.6(–1.8) mm, apex entire or emarginate with mucro to 0.3 mm.

Flowers

anthers (0.4–)0.6–1 mm;

styles 0.4–1.6 mm;

stigmas 1–1.5 mm.

anthers (0.2–)0.3–0.4(–0.7) mm;

styles 0.4–0.7(–1) mm;

stigmas (1–)1.5–3 mm.

Achenes

brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate.

brown, reddish brown, or black, stipitate, narrowly ellipsoid to oblong (rarely obovoid-oblong), (1–)1.2–1.5(–1.9) × 0.5–0.6(–0.75) mm, stipe 0.1–0.2 mm, apex acute to barely obtuse, surfaces finely papillose.

Cyperus hermaphroditus

Cyperus odoratus

Phenology Fruiting summer–fall. Fruiting summer–early fall.
Habitat Arroyos Emergent shorelines, disturbed, muddy places, fresh or slightly brackish marshes
Elevation 500–1200 m (1600–3900 ft) 0–1500 m (0–4900 ft)
Distribution
from FNA
AZ; TX; Mexico; South America; West Indies
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; HI; ON; QC; Central America; South America; Asia; Africa; Australia
[WildflowerSearch map]
[BONAP county map]
Discussion

An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

As circumscribed here, Cyperus odoratus is easily identified by its cylindric to subcylindric spikelets in which the corky rachilla of the mature spikelet disarticulates at the base of each scale. The mature spikelet breaks into segments each consisting of a scale and an internode of the rachilla clasping the achene with its corky wings. Cyperus odoratus is an exceedingly variable pantropical and warm-temperate species. Numerous segregates have been named, some of which may deserve recognition when the species is studied in detail worldwide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 186. FNA vol. 23, p. 191.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus Cyperaceae > Cyperus > subg. Diclidium
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii C. acicularis, C. californicus, C. eggersii, C. engelmannii, C. ferax, C. ferax subsp. engelmannii, C. ferax subsp. speciosus, C. ferruginescens, C. haenkei, C. huarmensis, C. macrocephalus, C. michauxianus, C. odoratus var. engelmannii, C. oxycarioides, C. speciosus, C. squarrosus var. parvus, Diclidium aciculare, Diclidium odoratum, Mariscus ferax, Mariscus huarmensis, Torulinium eggersii, Torulinium ferax, Torulinium macrocephalum, Torulinium michauxianum, Torulinium odoratum
Name authority (Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916) Linnaeus: Sp. Pl. 1: 46. (1753)
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