Cyperus hermaphroditus |
Cyperus niger |
|
---|---|---|
hermaphrodite flatsedge |
black flatsedge, brown cyperus |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, rhizomatous. |
Culms | trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous. |
solitary or close together, trigonous, 5–40(–95) cm, glabrous. |
Leaves | V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous. |
(2–)3–7, broadly V-shaped, 3–20(–30) cm × 1.5–3 mm. |
Inflorescences | spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide. |
spikes loosely to densely ovoid to hemispheric, 7–16 mm diam.; rachis 1–2 mm, or absent; rays 1–2, 3–45 mm; if rays absent, inflorescence sessile, dense, 6–20 mm diam; bracts 2–3, horizontal to reflexed downward, broadly V-shaped, 1–15 cm × 1–3 mm. |
Spikelets | (10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse. |
(3–)5–25(–60), linear to oblong-linear, (3–)5–9 × 1.8–2.3 mm; floral scales 4–18, closely imbricate, chestnut brown, black, or brown, medially green or greenish brown, laterally ribless, medially 2(–3)-ribbed, distinctly 2-keeled basally, ovate to orbiculate, 1.5–2.1 × 1.4–2.2 mm, apex obtuse. |
Flowers | anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm. |
stamens 2; anthers 0.6–0.8 mm, connectives not prolonged; styles 0.4–0.6 mm; stigmas 0.7–2.1 mm. |
Achenes | brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate. |
brown, ellipsoid, 1.2–1.4 × 0.6–0.8 mm, network of ridges forming isodiametric or square cells, slightly stipitate, apex apiculate, surfaces minutely punctate. |
Cyperus hermaphroditus |
Cyperus niger |
|
Phenology | Fruiting summer–fall. | Fruiting summer. |
Habitat | Arroyos | Swamps, ditches, wet pastures, seeps in montane forests |
Elevation | 500–1200 m (1600–3900 ft) | 0–1200 m (0–3900 ft) |
Distribution |
AZ; TX; Mexico; South America; West Indies |
AZ; CA; NM; OK; TX; Mexico; Central America; South America
|
Discussion | An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Six varieties (Cyperus niger var. niger, var. castaneus, var. robustus, var. lorentizianus, var. intricate-ramosus, and var. pseudo-elegantulus) were recognized by G. Kükenthal (1935–1936). Based on field observations and measurements of hundreds of collections from Mexico, where the species is most plentiful, it does not seem possible to recognize any of the infraspecific taxa (G. C. Tucker 1983, 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 186. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii | C. diandrus var. castaneus, C. melanostachyus, C. niger var. castaneus |
Name authority | (Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916) | Ruiz & Pavón: Fl. Peruv. 1: 47. (1798) |
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