FNA: Cyperus hermaphroditus vs. Cyperus lentiginosus

	Cyperus hermaphroditus	Cyperus lentiginosus
	hermaphrodite flatsedge	Latin American flatsedge
Habit	Herbs, perennial, rhizomatous.	Herbs, perennial, cespitose, rhizomatous.
Culms	trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous.	trigonous, (1–) 20–40(–75) cm × 1–1.5(–2.5) mm, glabrous.
Leaves	V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous.	inversely W-shaped, 10–40(–70) cm × (1–)3–6(–10) mm.
Inflorescences	spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide.	spikes 1(–3), broadly and loosely ovoid to oblong-ellipsoid, (15–)20–30(–50) × 20–30(–45) mm; bracts (3–)5–7(–10), ascending at 30(–45)°, inversely W-shaped, 10–30(–50) cm × (1–)3–5(–11) mm; rays (2–)3–6(–12), (1–)3–10(–17) cm; rachilla deciduous, wings ca. 0.3 mm wide.
Spikelets	(10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse.	(10–)25–50(–100), ± terete, (appearing compressed due to excurved apices of floral scales), (8–)12–15(–24) × (0.8–)1.1–1.4(–1.8) mm; floral scales persistent, (4–)6–8(–14), golden brown, off-white to stramineous, densely red-glandular punctate, green medially, 3–4-ribbed laterally, 3–5-ribbed medially, oblong-lanceolate, (3.2–)4–4.5 × 1.2–1.6 mm, apex blunt, mucronulate to mucronate, mucro 0.2–0.5 mm; proximal scales mucronulate, distal scales mucronate; terminal scale conduplicate.
Flowers	anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm.	anthers 0.5–0.8(–1.4) mm; styles 0.8–1 mm; stigmas 1–2 mm.
Achenes	brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate.	brown, ± sessile to stipitate, oblong-ellipsoid, 1.7–2 × 0.6–0.7 mm, 0.1(–0.2) × 0.2 mm, apex rounded, apiculate from dark purple style base, surfaces papillose.

	Cyperus hermaphroditus	Cyperus lentiginosus
Phenology	Fruiting summer–fall.	Fruiting spring–summer.
Habitat	Arroyos	Thickets, open woods
Elevation	500–1200 m (1600–3900 ft)	0–50 m (0–200 ft)
Distribution	AZ; TX; Mexico; South America; West Indies [BONAP county map]	FL; TX; Mexico; West Indies [BONAP county map]
Discussion	An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Cyperus lentiginosus has been treated as a variety of C. tenuis (G. Kükenthal 1935–1936). The two taxa differ in the longer scales with cuspidate apices of C. lentiginosus, the inversely W-shaped leaves and bracts (those of C. tenuis are V-shaped), and the open spikes (those of C. tenuis are dense). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23, p. 186.	FNA vol. 23.
Parent taxa	Cyperaceae > Cyperus > subg. Cyperus	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms	Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii	C. strigosus var. gracilis, C. tenuis var. lentiginosus
Name authority	(Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916)	Millspaugh & Chase: Publ. Field Columbian Mus., Bot. Ser. 3: 74. (1903)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus lentiginosus

hermaphrodite flatsedge

Latin American flatsedge

Habit

Herbs, perennial, rhizomatous.

Herbs, perennial, cespitose, rhizomatous.

Culms

trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous.

trigonous, (1–) 20–40(–75) cm × 1–1.5(–2.5) mm, glabrous.

Leaves

V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous.

inversely W-shaped, 10–40(–70) cm × (1–)3–6(–10) mm.

Inflorescences

spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis;

rays (3–)5–10(–17), (1–)5–12(–24) cm;

bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm;

rachilla deciduous, wings 0.4–0.5 mm wide.

spikes 1(–3), broadly and loosely ovoid to oblong-ellipsoid, (15–)20–30(–50) × 20–30(–45) mm;

bracts (3–)5–7(–10), ascending at 30(–45)°, inversely W-shaped, 10–30(–50) cm × (1–)3–5(–11) mm;

rays (2–)3–6(–12), (1–)3–10(–17) cm;

rachilla deciduous, wings ca. 0.3 mm wide.

Spikelets

(10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm;

distal spikelet spreading or ascending;

floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse.

(10–)25–50(–100), ± terete, (appearing compressed due to excurved apices of floral scales), (8–)12–15(–24) × (0.8–)1.1–1.4(–1.8) mm;

floral scales persistent, (4–)6–8(–14), golden brown, off-white to stramineous, densely red-glandular punctate, green medially, 3–4-ribbed laterally, 3–5-ribbed medially, oblong-lanceolate, (3.2–)4–4.5 × 1.2–1.6 mm, apex blunt, mucronulate to mucronate, mucro 0.2–0.5 mm;

proximal scales mucronulate, distal scales mucronate;

terminal scale conduplicate.

Flowers

anthers (0.4–)0.6–1 mm;

styles 0.4–1.6 mm;

stigmas 1–1.5 mm.

anthers 0.5–0.8(–1.4) mm;

styles 0.8–1 mm;

stigmas 1–2 mm.

Achenes

brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate.

brown, ± sessile to stipitate, oblong-ellipsoid, 1.7–2 × 0.6–0.7 mm, 0.1(–0.2) × 0.2 mm, apex rounded, apiculate from dark purple style base, surfaces papillose.

Cyperus hermaphroditus

Cyperus lentiginosus

Phenology

Fruiting summer–fall.

Fruiting spring–summer.

Habitat

Arroyos

Thickets, open woods

Elevation

500–1200 m (1600–3900 ft)

0–50 m (0–200 ft)

Distribution

AZ; TX; Mexico; South America; West Indies

[BONAP county map]

FL; TX; Mexico; West Indies

[BONAP county map]

Discussion

An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus lentiginosus has been treated as a variety of C. tenuis (G. Kükenthal 1935–1936). The two taxa differ in the longer scales with cuspidate apices of C. lentiginosus, the inversely W-shaped leaves and bracts (those of C. tenuis are V-shaped), and the open spikes (those of C. tenuis are dense).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23, p. 186.

FNA vol. 23.

Parent taxa

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii

C. strigosus var. gracilis, C. tenuis var. lentiginosus

Name authority

(Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916)

Millspaugh & Chase: Publ. Field Columbian Mus., Bot. Ser. 3: 74. (1903)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Cyperus hermaphroditus

Cyperus lentiginosus

Cyperus hermaphroditus

Cyperus lentiginosus