Cyperus hermaphroditus |
Cyperus laevigatus |
|
---|---|---|
hermaphrodite flatsedge |
smooth cyperus, smooth flatsedge, smooth nutgrass |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, rhizomatous. |
Culms | trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous. |
single or close together, trigonous to ± terete, (1–)8–30(–60) cm × (0.3–)0.6–1.5(–2.2) mm, glabrous. |
Leaves | V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous. |
blades present or reduced to sheaths, base of culm covered with reddish sheath 5–25 mm, distally short toothed 1–2 mm, or leaf blades involute, 1–7 cm × 0.3–1(–1.6) mm. |
Inflorescences | spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide. |
heads appearing lateral, spikes ± digitate, 5–15 × 5–15 mm; rays absent, sometimes 1, to 2 mm; bracts (1–)2(–3), longer bracts erect, appearing as continuation of culm, shorter bract(s) horizontal or ascending, longer bracts (1–)2–8(–12) cm × 0.5–3 mm, shorter (0.3–)1–4.5 cm × 0.2–1 mm. |
Spikelets | (10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse. |
(1–)2–7(–14), whitish green to reddish, compressed-turgid, ellipsoid to oblong-lanceoloid, 4–7(–12) × (1.6–)2–3(–3.6) mm; floral scales (8–)12–22, off-white, whitish with red speckles, or with single red spot 0.5 mm wide, medially whitish, oblong-laterally ribless or weakly 2-ribbed, medially weakly 1–3-ribbed, obovate to orbiculate, 1.5–2 × (1.3–)1.6–2 mm. |
Flowers | anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm. |
anthers (0.5–)0.7–1 mm, connective apex reddish, 0.1–0.2 mm; styles 0.8–1.2 mm; stigmas 0.7–1 mm. |
Achenes | brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate. |
gray, glossy, oblong-ellipsoid to ovoid, (1.2–)1.3–1.5(–1.8) × (0.7–)0.8–1(–1.2) mm, base cuneate to stipelike, apex apiculate, surfaces finely reticulate to papillate. |
Cyperus hermaphroditus |
Cyperus laevigatus |
|
Phenology | Fruiting summer–fall. | Fruiting summer. |
Habitat | Arroyos | Alkaline grasslands, hotsprings, ditches, brackish marshes |
Elevation | 500–1200 m (1600–3900 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AZ; TX; Mexico; South America; West Indies |
AZ; CA; NC; TX; Mexico; South America; West Indies; Europe; Asia; Africa
|
Discussion | An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus laevigatus is one of only two species of subg. Juncellus in the New World. It can be distinguished from all other New World species of Cyperus by its two stigmas and disclike, strongly flattened achene, which is borne with its face toward the rachilla. Species of subg. Pycreus, especially C. niger, have been misidentified as C. laevigatus, probably because of the dark, shiny spikelets. All species of subg. Pycreus have biconvex achenes like C. laevigatus; in species of subg. Pycreus the achene is attached with an edge toward the rachilla; in C. laevigatus the achene faces the rachilla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 186. | FNA vol. 23, p. 164. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Juncellus |
Sibling taxa | ||
Synonyms | Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii | Acorellus laevigatus, Chlorocyperus laevigatus, Juncellus laevigatus, Pycreus laevigatus |
Name authority | (Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916) | Linnaeus: Mant. Pl. 2: 179. (1771) |
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