FNA: Cyperus hermaphroditus vs. Cyperus fendlerianus

	Cyperus hermaphroditus	Cyperus fendlerianus
	hermaphrodite flatsedge	Fendler flatsedge, Fendler's flatsedge, Fendler's nutgrass, Fendler's sedge
Habit	Herbs, perennial, rhizomatous.	Herbs, perennial, rhizomatous.
Culms	trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous.	trigonous, (7–)20–70(–85) cm × (0.5–)1–1.8(–2.6) mm, glabrous or sparsely to densely scabridulous on surfaces or on angles immediately proximal to inflorescence in distal 3–6 cm.
Leaves	V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous.	2–3, flat to V-shaped, (12–)20–40(–55) cm × (2–)3–5(–7) mm.
Inflorescences	spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide.	spikes (1–)3–4(–6), largest erect, others at base of central spike, erect spike sessile, loosely to densely cylindric to ovoid-cylindric, (12–)18–30 × (6–)12–20 mm, remaining spikes 1/2 as large; rays absent; bracts (2–)3–6(–10), ± horizontal to reflexed parallel to culm, (2–)6–16(–25) cm × (1.5–)2–4(–7) mm; rachilla ± deciduous, wingless.
Spikelets	(10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse.	(8–)15–30, spreading to ascending-appressed, oblong-lanceoloid, quadrangular, slightly compressed, 5–8(–10) × (2–)2.4–3.4(–4.2) mm; floral scales (2–)4–6(–8), yellowish, reddish, or greenish brown, medially greenish, straight to excurved, laterally (2–)3–4(–5)-ribbed, medially (1–)3–(–5)-ribbed, ovate-orbiculate, 2.4–2.8(–3.4) × 2.4–2.8(–3.4) mm, apex obtuse, mucronate to cuspidate, stout, (0.1–)0.3–0.6 mm, medially glabrous.
Flowers	anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm.	anthers 0.4–0.8 mm; styles 0.7–1.3 mm; stigmas 0.7–1.2 mm.
Achenes	brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate.	finely reticulate, brown to reddish brown, ± stipelike, obovoid, 1.6–1.9 × 1.1–1.4(–1.6) mm, base attenuate to cuneate, 0.1–0.2 × 0.2 mm, apex obtuse, apiculate, surfaces glabrous.

	Cyperus hermaphroditus	Cyperus fendlerianus
Phenology	Fruiting summer–fall.	Fruiting summer.
Habitat	Arroyos	Clearings in montane forests
Elevation	500–1200 m (1600–3900 ft)	500–1800 m (1600–5900 ft)
Distribution	AZ; TX; Mexico; South America; West Indies [BONAP county map]	AZ; CO; NM; TX; WY; Mexico [WildflowerSearch map] [BONAP county map]
Discussion	An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Hybridization between Cyperus fendlerianus and C. sphaerolepis occurs occasionally in the southwestern United States and northern Mexico (B. G. Marcks 1972). The unnamed hybrid appears to be fully fertile and can be recognized by morphologic intermediacy. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23, p. 186.	FNA vol. 23, p. 177.
Parent taxa	Cyperaceae > Cyperus > subg. Cyperus	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms	Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii	Chlorocyperus fendlerianus
Name authority	(Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916)	Boeckeler: Linnaea 35: 520. (1868)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus fendlerianus

hermaphrodite flatsedge

Fendler flatsedge, Fendler's flatsedge, Fendler's nutgrass, Fendler's sedge

Habit

Herbs, perennial, rhizomatous.

Culms

trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous.

trigonous, (7–)20–70(–85) cm × (0.5–)1–1.8(–2.6) mm, glabrous or sparsely to densely scabridulous on surfaces or on angles immediately proximal to inflorescence in distal 3–6 cm.

Leaves

V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous.

2–3, flat to V-shaped, (12–)20–40(–55) cm × (2–)3–5(–7) mm.

Inflorescences

spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis;

rays (3–)5–10(–17), (1–)5–12(–24) cm;

bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm;

rachilla deciduous, wings 0.4–0.5 mm wide.

spikes (1–)3–4(–6), largest erect, others at base of central spike, erect spike sessile, loosely to densely cylindric to ovoid-cylindric, (12–)18–30 × (6–)12–20 mm, remaining spikes 1/2 as large;

rays absent;

bracts (2–)3–6(–10), ± horizontal to reflexed parallel to culm, (2–)6–16(–25) cm × (1.5–)2–4(–7) mm;

rachilla ± deciduous, wingless.

Spikelets

(10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm;

distal spikelet spreading or ascending;

floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse.

(8–)15–30, spreading to ascending-appressed, oblong-lanceoloid, quadrangular, slightly compressed, 5–8(–10) × (2–)2.4–3.4(–4.2) mm;

floral scales (2–)4–6(–8), yellowish, reddish, or greenish brown, medially greenish, straight to excurved, laterally (2–)3–4(–5)-ribbed, medially (1–)3–(–5)-ribbed, ovate-orbiculate, 2.4–2.8(–3.4) × 2.4–2.8(–3.4) mm, apex obtuse, mucronate to cuspidate, stout, (0.1–)0.3–0.6 mm, medially glabrous.

Flowers

anthers (0.4–)0.6–1 mm;

styles 0.4–1.6 mm;

stigmas 1–1.5 mm.

anthers 0.4–0.8 mm;

styles 0.7–1.3 mm;

stigmas 0.7–1.2 mm.

Achenes

brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate.

finely reticulate, brown to reddish brown, ± stipelike, obovoid, 1.6–1.9 × 1.1–1.4(–1.6) mm, base attenuate to cuneate, 0.1–0.2 × 0.2 mm, apex obtuse, apiculate, surfaces glabrous.

Cyperus hermaphroditus

Cyperus fendlerianus

Phenology

Fruiting summer–fall.

Fruiting summer.

Habitat

Arroyos

Clearings in montane forests

Elevation

500–1200 m (1600–3900 ft)

500–1800 m (1600–5900 ft)

Distribution

AZ; TX; Mexico; South America; West Indies

[BONAP county map]

AZ; CO; NM; TX; WY; Mexico

[WildflowerSearch map]

[BONAP county map]

Discussion

An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybridization between Cyperus fendlerianus and C. sphaerolepis occurs occasionally in the southwestern United States and northern Mexico (B. G. Marcks 1972). The unnamed hybrid appears to be fully fertile and can be recognized by morphologic intermediacy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23, p. 186.

FNA vol. 23, p. 177.

Parent taxa

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii

Chlorocyperus fendlerianus

Name authority

(Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916)

Boeckeler: Linnaea 35: 520. (1868)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Cyperus hermaphroditus

Cyperus fendlerianus

Cyperus hermaphroditus

Cyperus fendlerianus