Cyperus hermaphroditus |
Cyperus entrerianus |
|
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hermaphrodite flatsedge |
woodrush flatsedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. |
Culms | trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous. |
1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
Leaves | V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous. |
(1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
Inflorescences | spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide. |
heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
Spikelets | (10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse. |
(1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
Flowers | anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm. |
stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
Achenes | brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate. |
brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
Cyperus hermaphroditus |
Cyperus entrerianus |
|
Phenology | Fruiting summer–fall. | Fruiting summer. |
Habitat | Arroyos | Roadside ditches, marshes |
Elevation | 500–1200 m (1600–3900 ft) | 0–100 m (0–300 ft) |
Distribution |
AZ; TX; Mexico; South America; West Indies |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
|
Discussion | An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 186. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii | C. luzulae var. entrerianus |
Name authority | (Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916) | Boeckeler: Flora 61: 139. (1878) |
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