Cyperus hermaphroditus |
Cyperus drummondii |
|
---|---|---|
hermaphrodite flatsedge |
Drummond's sedge |
|
Habit | Herbs, perennial, rhizomatous. | Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. |
Culms | trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous. |
trigonous, 35–170 cm × 2–4.7 mm, scabrid on angles. |
Leaves | V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous. |
4–12, with readily visible cross ribs especially on abaxial surface, flat to V-shaped, 25–100 cm × 2.4–11 mm. |
Inflorescences | spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide. |
heads digitate, globose, 1–1.7 cm diam.; rays 3–5, 1–7 cm; 2d order rays absent; bracts 3–5, horizontal to ascending at 30°, with prominent cross ribs, V-shaped, (4–)11–35(–48) cm × 1–6.7 mm (longest 1/2 as long as culm); 2d order bracts absent. |
Spikelets | (10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse. |
10–40, oblong to linear-lanceoloid, 5–1.8 × 1.5–2.2 mm; floral scales 18–42, pale grayish green, turning brownish, proximally greenish or brownish, 2-keeled, weakly to distinctly 1-ribbed, proximal 1/2 2-ribbed, ovate, 1.4–1.6 × 0.9–1.4 mm, acute (sometimes mucronulate), apically glabrous, occasionally scabridulous. |
Flowers | anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm. |
stamens 1–2; anthers 0.8–1.2 mm; styles 0.8–1 mm; stigmas 0.6–0.8 mm. |
Achenes | brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate. |
brown, slightly stipitate, oblong-ellipsoid (2.4–3.9 times longer than wide), 1–1.2 × 0.3–0.5 mm, apical beak 0.1–0.5 mm, surfaces glabrous. |
Cyperus hermaphroditus |
Cyperus drummondii |
|
Phenology | Fruiting summer–fall. | Fruiting late spring–summer. |
Habitat | Arroyos | Wet, relatively undisturbed habitats, especially flatwoods, ponds, seepage slopes, coastal prairies |
Elevation | 500–1200 m (1600–3900 ft) | 0–200 m (0–700 ft) |
Distribution |
AZ; TX; Mexico; South America; West Indies |
AL; FL; GA; LA; MS; SC; TX; Central America; South America; West Indies |
Discussion | An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus drummondii has been treated as a synonym (G. C. Tucker 1994), or a variety (G. Kükenthal 1935–1936), or a subspecies of C. virens (M. F. Denton 1978b). Recent quantitative and field studies (J. R. Carter et al. 1999) provided compelling evidence that specific status is appropriate. Compared to Cyperus virens, C. drummondii is a taller species of less disturbed habitats and has narrower spikelets, ovate scales, and fewer inflorescence bracts. Cyperus virens and C. drummondii are readily distinguished from other Cyperus species in the United States by their sharply angled, scabrid culms and conspicuously septate leaf blades. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 186. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii | C. robustus, C. virens subsp. drummondii, C. virens var. drummondii, C. virens var. robustus |
Name authority | (Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916) | Torrey & Hooker: Ann. Lyceum Nat. Hist. New York 3: 437, errata. (1836) |
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