FNA: Cyperus hermaphroditus vs. Cyperus dentoniae

	Cyperus hermaphroditus	Cyperus dentoniae
	hermaphrodite flatsedge	Denton's flatsedge, hairy flatsedge
Habit	Herbs, perennial, rhizomatous.	Herbs, perennial, cespitose, shortly rhizomatous.
Culms	trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous.	1–4, trigonous, (2–)20–50(–80) cm × (0.8–)1.5–2(–2.8) mm, basally glabrous or nearly so, apically hispidulous, on faces and angles distally, especially so immediately proximal to bracts.
Leaves	V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous.	2–5(–7), V-shaped, (5–)15–30(–45) cm × (1.5–)2.5–5(–8) mm.
Inflorescences	spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide.	spike 1 (occasionally 1–2 smaller sessile spikes at base), densely cylindric, (7–)11–20(–25) × (5–) 7–10(–12) mm; rays 3–6(–10), (1–)3–6(–18) cm, rays and rachis hispidulous; bracts 4–8(–10), ascending at 30–60°, V-shaped, (2–)10–25(–45) cm × (1–)3–6(–8) mm; rachilla deciduous, wings persistent, 0.3–0.4 mm wide.
Spikelets	(10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse.	(20–)70–120(–140), oblong-ellipsoid to oblong-lanceoloid, ± terete to quadrangular, (2.2–)3.4–4.5(–5.6) × (0.8–)0.9–1.2(–1.3) mm; distal spikelet spreading or ascending; floral scales 1(–3), appressed, stramineous, red-spotted, strongly 3(–4)-ribbed on either side of green part, medially weakly 3-ribbed, ovate to elliptic ovate, (2.4–)2.8–3.3(–3.6) × 1.6–2 mm, apex acute, mucronulate from excurrent midrib.
Flowers	anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm.	anthers (0.3–)0.4–0.6 mm; styles 0.4–0.6(–0.8) mm; stigmas 1–1.4 mm.
Achenes	brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate.	brown, stipitate, ellipsoid, (1.6–)1.8–2 × (0.7–) 0.8–0.9 mm, stipe 0.1–0.2 × 0.2 mm, apex ± truncate, apiculate, surfaces puncticulate to glabrous.

	Cyperus hermaphroditus	Cyperus dentoniae
Phenology	Fruiting summer–fall.	Fruiting mid summer–fall (Jul–Sep).
Habitat	Arroyos	Various dry, open environments
Elevation	500–1200 m (1600–3900 ft)	500–1000 m (1600–3300 ft)
Distribution	AZ; TX; Mexico; South America; West Indies [BONAP county map]	AZ; Mexico; Central America; South America [BONAP county map]
Discussion	An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Cyperus dentoniae was not treated in G. Kükenthal’s monograph (1935–1936), apparently because he did not see any materials of the species. The plant has been treated as “C. asper (Liebmann) O’Neill,” a name based on Mariscus asper Liebmann and a synonym of C. mutisii (A. B. Ayers 1946). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23, p. 186.	FNA vol. 23, p. 187.
Parent taxa	Cyperaceae > Cyperus > subg. Cyperus	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms	Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii	Mariscus pubescens, C. flavomariscus var. peduncularis, C. flavus var. peduncularis
Name authority	(Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916)	G. C. Tucker: Syst. Bot. Monogr. 2: 56. (1983)
Web links	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus dentoniae

hermaphrodite flatsedge

Denton's flatsedge, hairy flatsedge

Habit

Herbs, perennial, rhizomatous.

Herbs, perennial, cespitose, shortly rhizomatous.

Culms

trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous.

1–4, trigonous, (2–)20–50(–80) cm × (0.8–)1.5–2(–2.8) mm, basally glabrous or nearly so, apically hispidulous, on faces and angles distally, especially so immediately proximal to bracts.

Leaves

V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous.

2–5(–7), V-shaped, (5–)15–30(–45) cm × (1.5–)2.5–5(–8) mm.

Inflorescences

spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis;

rays (3–)5–10(–17), (1–)5–12(–24) cm;

bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm;

rachilla deciduous, wings 0.4–0.5 mm wide.

spike 1 (occasionally 1–2 smaller sessile spikes at base), densely cylindric, (7–)11–20(–25) × (5–) 7–10(–12) mm;

rays 3–6(–10), (1–)3–6(–18) cm, rays and rachis hispidulous;

bracts 4–8(–10), ascending at 30–60°, V-shaped, (2–)10–25(–45) cm × (1–)3–6(–8) mm;

rachilla deciduous, wings persistent, 0.3–0.4 mm wide.

Spikelets

(10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm;

distal spikelet spreading or ascending;

floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse.

(20–)70–120(–140), oblong-ellipsoid to oblong-lanceoloid, ± terete to quadrangular, (2.2–)3.4–4.5(–5.6) × (0.8–)0.9–1.2(–1.3) mm;

distal spikelet spreading or ascending;

floral scales 1(–3), appressed, stramineous, red-spotted, strongly 3(–4)-ribbed on either side of green part, medially weakly 3-ribbed, ovate to elliptic ovate, (2.4–)2.8–3.3(–3.6) × 1.6–2 mm, apex acute, mucronulate from excurrent midrib.

Flowers

anthers (0.4–)0.6–1 mm;

styles 0.4–1.6 mm;

stigmas 1–1.5 mm.

anthers (0.3–)0.4–0.6 mm;

styles 0.4–0.6(–0.8) mm;

stigmas 1–1.4 mm.

Achenes

brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate.

brown, stipitate, ellipsoid, (1.6–)1.8–2 × (0.7–) 0.8–0.9 mm, stipe 0.1–0.2 × 0.2 mm, apex ± truncate, apiculate, surfaces puncticulate to glabrous.

Cyperus hermaphroditus

Cyperus dentoniae

Phenology

Fruiting summer–fall.

Fruiting mid summer–fall (Jul–Sep).

Habitat

Arroyos

Various dry, open environments

Elevation

500–1200 m (1600–3900 ft)

500–1000 m (1600–3300 ft)

Distribution

AZ; TX; Mexico; South America; West Indies

[BONAP county map]

AZ; Mexico; Central America; South America

[BONAP county map]

Discussion

An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus dentoniae was not treated in G. Kükenthal’s monograph (1935–1936), apparently because he did not see any materials of the species. The plant has been treated as “C. asper (Liebmann) O’Neill,” a name based on Mariscus asper Liebmann and a synonym of C. mutisii (A. B. Ayers 1946).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23, p. 186.

FNA vol. 23, p. 187.

Parent taxa

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii

Mariscus pubescens, C. flavomariscus var. peduncularis, C. flavus var. peduncularis

Name authority

(Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916)

G. C. Tucker: Syst. Bot. Monogr. 2: 56. (1983)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Cyperus hermaphroditus

Cyperus dentoniae

Cyperus hermaphroditus

Cyperus dentoniae