Cyperus hermaphroditus |
Cyperus |
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hermaphrodite flatsedge |
cyperus, flatsedge, galingale, nutsedge, umbrella-sedge |
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Habit | Herbs, perennial, rhizomatous. | Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous. | ||||||||||||||||
Culms | trigonous, (10–)30–80(–140) cm × (0.4–)1–3(–5) mm, glabrous. |
solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles. |
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Leaves | V-shaped or flanged V-shaped, (5–)15–70(–90) cm × (2–)4–10(–13) mm, glabrous. |
usually basal; ligules absent; blades keeled abaxially, flat, V-, or inversely W-shaped in cross section. |
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Inflorescences | spikes 1(–5), rather densely ovoid or oblong-cylindric, (10–)25–45(–62) × 7–22 mm, 11–21 spikelets per 5 mm of rachis; rays (3–)5–10(–17), (1–)5–12(–24) cm; bracts (2–)4–8, ascending at 30–45°, V- or inversely W-shaped, (2–)8–40(–65) cm × (1–)2–8 mm; rachilla deciduous, wings 0.4–0.5 mm wide. |
terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet; spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts; involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike. |
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Spikelets | (10–)50–150, oblong to linear, quadrangular, (3–)4–8(–18) × (0.8–)1–1.2(–1.6) mm; distal spikelet spreading or ascending; floral scales (1–) 3–8, pale greenish white to golden brown, deep brown, or reddish brown, medially usually green, weakly to strongly 3(–4)-ribbed laterally, 1–3-ribbed medially, ovate to elliptic, (2–)2.6–3.2(–4) × 1.2–2 mm, apex obtuse. |
scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous). |
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Flowers | anthers (0.4–)0.6–1 mm; styles 0.4–1.6 mm; stigmas 1–1.5 mm. |
bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings; perianth absent; stamens 1–3; styles linear, 2–3-fid, base deciduous or persistent; stigmas 2–3. |
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Achenes | brown or reddish brown, sessile or nearly so, ellipsoid, 1.4–1.8(–2) × 0.6–0.8(–1) mm, apex acute to obtuse, slightly apiculate, surfaces puncticulate. |
biconvex, flattened, or trigonous. |
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Cyperus hermaphroditus |
Cyperus |
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Phenology | Fruiting summer–fall. | |||||||||||||||||
Habitat | Arroyos | |||||||||||||||||
Elevation | 500–1200 m (1600–3900 ft) | |||||||||||||||||
Distribution |
AZ; TX; Mexico; South America; West Indies |
Pantemperate and tropical |
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Discussion | An erroneous report of Cyperus hermaphroditus from Alabama was based on misidentification of C. thyrsiflorus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 600 (96 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 186. | FNA vol. 23, p. 141. | ||||||||||||||||
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae | ||||||||||||||||
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Synonyms | Carex hermaphrodita, Mariscus hermaphroditus, Mariscus jacquinii | |||||||||||||||||
Name authority | (Jacquin) Standley: Contr. U.S. Natl. Herb. 18: 88. (1916) | Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754) | ||||||||||||||||
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