Cyperus haspan |
Cyperus reflexus |
|
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haspan flatsedge |
bentawn flatsedge |
|
Habit | Herbs, perennial, cespitose, shortly rhizomatous. | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. |
Culms | trigonous, (2–)25–60(–100) cm × 1–4 mm, soft (flattened in drying), glabrous. |
roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
Leaves | usually reduced to sheaths, occasionally with blades, flat to V-shaped, (3–)10–30 cm × (1–)2.5–5 mm. |
1–6, 10–40 cm, glabrous or essentially so. |
Inflorescences | heads loosely digitate; rays (5–)10–12(–15), 1–15(–20) cm; 2d order rays usually present, (1–)6–12(–25) mm; 3d order rays sometimes present, 1–6 mm; bracts 2(–3), horizontal to ascending at 30–60°, longer (3–)6–18 cm × 2–4 mm, shorter (0.5–)2.5–6.5 cm × 1–2(–3) mm. |
heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
Spikelets | 1–15, linear-lanceoloid, ± compressed-quadrangular, 3–18 × 1–1.6 mm; floral scales 10–20(–40), laterally reddish to greenish brown, dull, often clear-edged, medially greenish, laterally 1-ribbed, medially 1-ribbed, oblong to obovate, 1–1.5(–1.9) × 0.8–1 mm, apex mucronate, glabrous except for cluster of crystalline prickles at apex. |
25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
Flowers | stamens 3; anthers 0.3–0.5 mm; styles 0.4–0.9 mm; stigmas 0.5–0.8(–1.3) mm. |
stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
Achenes | white or reddish brown, stipitate, globose to obovoid or ellipsoid, 0.5–0.6(–0.7) × 0.3–0.4(–0.5) mm, base nearly cuneate, stipe 0.1 × 0.1–0.2 mm, apex obtuse to acute, apiculate or entire, surfaces granular to papillose. |
reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
Cyperus haspan |
Cyperus reflexus |
|
Phenology | Fruiting summer. | Fruiting late spring–fall (May–Oct). |
Habitat | Swales between dunes, marshes, pond shores | Muddy soils or shallow waters |
Elevation | 0–200 m (0–700 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; FL; GA; LA; MS; NC; SC; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
Discussion | Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 23, p. 151. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | C. autumnalis, C. haspan subsp. juncoides, C. haspan var. americanus, C. juncoides | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus |
Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Vahl: Enum. Pl. 2: 299. (1805) |
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