Cyperus haspan |
Cyperus polystachyos |
|
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haspan flatsedge |
many-spike flatsedge |
|
Habit | Herbs, perennial, cespitose, shortly rhizomatous. | Herbs, perennial (annual?), cespitose. |
Culms | trigonous, (2–)25–60(–100) cm × 1–4 mm, soft (flattened in drying), glabrous. |
trigonous or ± terete, (3–)20–45(–70) cm × (0.8–) 1.2–2 mm, glabrous. |
Leaves | usually reduced to sheaths, occasionally with blades, flat to V-shaped, (3–)10–30 cm × (1–)2.5–5 mm. |
(2–) 6–10(–16), flat to V-shaped, (2–) 10–30(–60) cm × 1–4 mm. |
Inflorescences | heads loosely digitate; rays (5–)10–12(–15), 1–15(–20) cm; 2d order rays usually present, (1–)6–12(–25) mm; 3d order rays sometimes present, 1–6 mm; bracts 2(–3), horizontal to ascending at 30–60°, longer (3–)6–18 cm × 2–4 mm, shorter (0.5–)2.5–6.5 cm × 1–2(–3) mm. |
spike 1, loosely ovoid to densely turbinate, 12–40 × 10–25 mm; rachis 3–10 mm; rays 1–8(–12), 0.5–3(–6) cm; 2d order rays absent; if rays absent, inflorescence single dense cluster of spikelets, 12–40 mm diam.; bracts (3–)4–6(–7), horizontal to ascending at 45°, flanged V-shaped, (2–)5–15(–23) cm × 1–4 mm. |
Spikelets | 1–15, linear-lanceoloid, ± compressed-quadrangular, 3–18 × 1–1.6 mm; floral scales 10–20(–40), laterally reddish to greenish brown, dull, often clear-edged, medially greenish, laterally 1-ribbed, medially 1-ribbed, oblong to obovate, 1–1.5(–1.9) × 0.8–1 mm, apex mucronate, glabrous except for cluster of crystalline prickles at apex. |
(5–)15–30(–40), spreading to appressed-ascending, linear-lanceoloid to linear, compressed, 8–18(–40) × (1–)1.3–1.6(–2) mm; floral scales (8–)12–25(–66), laterally stramineous to light brown, medially green, laterally ribless, medially 3–5-ribbed, oblong, (1.5–)1.8–2.4 × (1–)1.2–1.4 mm, apex acute to ± acute, mucronate. |
Flowers | stamens 3; anthers 0.3–0.5 mm; styles 0.4–0.9 mm; stigmas 0.5–0.8(–1.3) mm. |
stamens 2; anthers (0.4–)0.5–0.8 mm, connective not prolonged; styles 0.6–1(–1.4) mm; stigmas (1–)1.4–2 mm. |
Achenes | white or reddish brown, stipitate, globose to obovoid or ellipsoid, 0.5–0.6(–0.7) × 0.3–0.4(–0.5) mm, base nearly cuneate, stipe 0.1 × 0.1–0.2 mm, apex obtuse to acute, apiculate or entire, surfaces granular to papillose. |
light to dark brown, oblong, (0.8–)0.9–1.1(–1.2) × 0.4–0.5(–0.6) mm, base cuneate to ± stipelike, apex truncate, apiculate, surfaces punctate to minutely punctate with raised cells. |
Cyperus haspan |
Cyperus polystachyos |
|
Phenology | Fruiting summer. | Fruiting summer–early fall. |
Habitat | Swales between dunes, marshes, pond shores | Shores, ditches, swales between dunes |
Elevation | 0–200 m [0–700 ft] | 0–200 m [0–700 ft] |
Distribution |
AL; FL; GA; LA; MS; NC; SC; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
|
AL; DC; DE; FL; GA; KY; LA; MA; MD; ME; MS; NC; NJ; NY; OK; PA; SC; TX; VA; WV; Mexico; Central America; South America; Asia; Africa
|
Discussion | Cyperus polystachyos is polymorphic. Adequate understanding of the infraspecific variations of C. polystachyos requires a worldwide examination, which is beyond the scope of this study. The segregates, C. fugax and C. filicinus, are recognized here; they have been usually accepted by recent workers (e.g., M. L. Fernald 1950; H. A. Gleason and A. Cronquist 1991). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 23, p. 151. | FNA vol. 23, p. 160. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. autumnalis, C. haspan subsp. juncoides, C. haspan var. americanus, C. juncoides | C. brizaeus, C. polystachyos var. leptostachyus, Pycreus polystachyos |
Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Rottbøll: Descr. Pl. Rar., 21. (1772) |
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