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haspan flatsedge

granite flatsedge

Habit Herbs, perennial, cespitose, shortly rhizomatous. Herbs, annual, cespitose, with roots fibrous.
Culms

trigonous, (2–)25–60(–100) cm × 1–4 mm, soft (flattened in drying), glabrous.

trigonous, glabrous.

Leaves

usually reduced to sheaths, occasionally with blades, flat to V-shaped, (3–)10–30 cm × (1–)2.5–5 mm.

V-shaped to flat, (2–)4–11 cm × 3–4 mm.

Inflorescences

heads loosely digitate;

rays (5–)10–12(–15), 1–15(–20) cm; 2d order rays usually present, (1–)6–12(–25) mm; 3d order rays sometimes present, 1–6 mm;

bracts 2(–3), horizontal to ascending at 30–60°, longer (3–)6–18 cm × 2–4 mm, shorter (0.5–)2.5–6.5 cm × 1–2(–3) mm.

rays absent;

bracts 2–4, ± horizontal, flat, 1–6.5 cm × 2–3 mm;

rachilla persistent, wingless.

Spikelets

1–15, linear-lanceoloid, ± compressed-quadrangular, 3–18 × 1–1.6 mm;

floral scales 10–20(–40), laterally reddish to greenish brown, dull, often clear-edged, medially greenish, laterally 1-ribbed, medially 1-ribbed, oblong to obovate, 1–1.5(–1.9) × 0.8–1 mm, apex mucronate, glabrous except for cluster of crystalline prickles at apex.

10–40, compressed, ovoid to linear-lanceoloid, quadrangular, 5–9 × 2–3 mm;

floral scales deciduous, 5–20, spreading, pale greenish to reddish brown, laterally (5–)7–9(–13)-ribbed, ovate, 3–4 × 1–2 mm, apex straight, excurved mucronate, or cuspidate.

Flowers

stamens 3;

anthers 0.3–0.5 mm;

styles 0.4–0.9 mm;

stigmas 0.5–0.8(–1.3) mm.

stamens 1–2;

filaments 1.5 mm;

anthers 0.8–1 mm, connective apex reddish yellow, less than 0.1 mm;

styles 0.8–1 mm;

stigmas 1–1.2 mm.

Achenes

white or reddish brown, stipitate, globose to obovoid or ellipsoid, 0.5–0.6(–0.7) × 0.3–0.4(–0.5) mm, base nearly cuneate, stipe 0.1 × 0.1–0.2 mm, apex obtuse to acute, apiculate or entire, surfaces granular to papillose.

brown to reddish brown, broadly ellipsoid to obovoid, 0.8–1.2 × 0.5–0.6 mm, base ± cuneate, apex truncate-obtuse, surfaces puncticulate.

Cyperus haspan

Cyperus granitophilus

Phenology Fruiting summer. Fruiting summer.
Habitat Swales between dunes, marshes, pond shores Thin soil over outcrops of granite or sandstone of the Piedmont and inner coastal plain
Elevation 0–200 m (0–700 ft) 50–400 m (200–1300 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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from FNA
AL; GA; NC; SC; TN; VA
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[BONAP county map]
Discussion

Cyperus granitophilus is documented as an autotetraploid derivative of C. squarrosus (L. W. Garoni and W. H. Murdy 1964).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 151. FNA vol. 23, p. 165.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. autumnalis, C. haspan subsp. juncoides, C. haspan var. americanus, C. juncoides
Name authority Linnaeus: Sp. Pl. 1: 45. (1753) McVaugh: Castanea 2: 103, figs. 4–8. (1937)
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