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haspan flatsedge

tooth flatsedge

Habit Herbs, perennial, cespitose, shortly rhizomatous. Herbs, perennial, cespitose, rhizomatous; stolons tuberiferous, 1–10 cm.
Culms

trigonous, (2–)25–60(–100) cm × 1–4 mm, soft (flattened in drying), glabrous.

trigonous, 8–50 cm × 1–3 mm.

Leaves

usually reduced to sheaths, occasionally with blades, flat to V-shaped, (3–)10–30 cm × (1–)2.5–5 mm.

flat or V-shaped, 10–40 cm × 2–5 mm.

Inflorescences

heads loosely digitate;

rays (5–)10–12(–15), 1–15(–20) cm; 2d order rays usually present, (1–)6–12(–25) mm; 3d order rays sometimes present, 1–6 mm;

bracts 2(–3), horizontal to ascending at 30–60°, longer (3–)6–18 cm × 2–4 mm, shorter (0.5–)2.5–6.5 cm × 1–2(–3) mm.

heads digitate, 8–18 mm diam.;

rays 4–9, 1–8 cm; 2d order rays 0.5–2 cm;

bracts 3–5, ± horizontal, flat, 1–20 cm × 1–4 mm.

Spikelets

1–15, linear-lanceoloid, ± compressed-quadrangular, 3–18 × 1–1.6 mm;

floral scales 10–20(–40), laterally reddish to greenish brown, dull, often clear-edged, medially greenish, laterally 1-ribbed, medially 1-ribbed, oblong to obovate, 1–1.5(–1.9) × 0.8–1 mm, apex mucronate, glabrous except for cluster of crystalline prickles at apex.

commonly proliferous, 2–6, linear to oblong, compressed, 3–24 × 2–2.5 mm;

floral scales 3–20(–50), reddish brown, laterally 1–3-ribbed, ovate, 2–2.5 × 1.5–2 mm.

Flowers

stamens 3;

anthers 0.3–0.5 mm;

styles 0.4–0.9 mm;

stigmas 0.5–0.8(–1.3) mm.

stamens 3;

anthers 1.5 mm;

styles 1 mm;

stigmas 1.5 mm.

Achenes

white or reddish brown, stipitate, globose to obovoid or ellipsoid, 0.5–0.6(–0.7) × 0.3–0.4(–0.5) mm, base nearly cuneate, stipe 0.1 × 0.1–0.2 mm, apex obtuse to acute, apiculate or entire, surfaces granular to papillose.

reddish brown, sessile, obovoid, 0.8–1 × 0.4–0.5 mm, apex obtuse, scarcely apiculate, surfaces glabrous.

Cyperus haspan

Cyperus dentatus

Phenology Fruiting summer. Fruiting summer.
Habitat Swales between dunes, marshes, pond shores Emergent sandy, peaty, or gravelly shorelines
Elevation 0–200 m (0–700 ft) 0–600 m (0–2000 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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from FNA
CT; DC; DE; IN; MA; MD; ME; NC; NH; NY; PA; RI; SC; TN; VA; VT; NB; NS; ON; QC
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Discussion

The report of Cyperus dentatus from West Virginia (M. V. McGivney 1938) is erroneous; the cited specimen was from New Jersey (Gutenberg s.n. 1886, US) (M. Strong, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 151. FNA vol. 23, p. 151.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Pycnostachys
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. autumnalis, C. haspan subsp. juncoides, C. haspan var. americanus, C. juncoides
Name authority Linnaeus: Sp. Pl. 1: 45. (1753) Torrey: Fl. N. Middle United States 1: 61. (1824)
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