Cyperus haspan |
Cyperus cephalanthus |
|
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haspan flatsedge |
buttonbush flatsedge |
|
Habit | Herbs, perennial, cespitose, shortly rhizomatous. | Herbs, perennial, rhizomatous. |
Culms | trigonous, (2–)25–60(–100) cm × 1–4 mm, soft (flattened in drying), glabrous. |
trigonous, 40–70(–100) cm × 2–3 mm, scabrid or hirtellate, either immediately proximal to apex or over distal 1/2 of culm. |
Leaves | usually reduced to sheaths, occasionally with blades, flat to V-shaped, (3–)10–30 cm × (1–)2.5–5 mm. |
flat, 30–80 cm × 4–10 mm, scabrid on margins, ribs on abaxial surface. |
Inflorescences | heads loosely digitate; rays (5–)10–12(–15), 1–15(–20) cm; 2d order rays usually present, (1–)6–12(–25) mm; 3d order rays sometimes present, 1–6 mm; bracts 2(–3), horizontal to ascending at 30–60°, longer (3–)6–18 cm × 2–4 mm, shorter (0.5–)2.5–6.5 cm × 1–2(–3) mm. |
spikes ovoid, 12–20 × 13–18 mm; rays 3–6, 4–10(–16) cm, glabrous; bracts 3–5, ± horizontal to reflexed, 3–12(–36) cm × 0.4–3(–5) mm, scabrid like leaves; rachilla deciduous, essentially wingless or wings inconspicuous, hyaline, 0.1(–0.2) mm wide. |
Spikelets | 1–15, linear-lanceoloid, ± compressed-quadrangular, 3–18 × 1–1.6 mm; floral scales 10–20(–40), laterally reddish to greenish brown, dull, often clear-edged, medially greenish, laterally 1-ribbed, medially 1-ribbed, oblong to obovate, 1–1.5(–1.9) × 0.8–1 mm, apex mucronate, glabrous except for cluster of crystalline prickles at apex. |
25–50(–80), oblong, quadrangular-compressed, 5–10 × 2.5–3.5 mm; floral scales deciduous, 4–10(–14), marginally clear, laterally brownish to clear distally, or light brown or reddish brown, blunt, laterally 3(–4)-ribbed, ovate-lanceolate, 2.5–3 × (1.2–)1.4–2 mm, often erose apically, apex acute to obtuse, entire or emarginate. |
Flowers | stamens 3; anthers 0.3–0.5 mm; styles 0.4–0.9 mm; stigmas 0.5–0.8(–1.3) mm. |
anthers 0.4–0.6 mm; styles 0.8–1.2 mm; stigmas 2 mm. |
Achenes | white or reddish brown, stipitate, globose to obovoid or ellipsoid, 0.5–0.6(–0.7) × 0.3–0.4(–0.5) mm, base nearly cuneate, stipe 0.1 × 0.1–0.2 mm, apex obtuse to acute, apiculate or entire, surfaces granular to papillose. |
brown to reddish brown, ± stipitate, ellipsoid, 1.2 × 0.5–0.6 mm, apex obtuse, scarcely to distinctly apiculate, surfaces puncticulate. |
Cyperus haspan |
Cyperus cephalanthus |
|
Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Swales between dunes, marshes, pond shores | Wet prairies |
Elevation | 0–200 m (0–700 ft) | 0–50 m (0–200 ft) |
Distribution |
AL; FL; GA; LA; MS; NC; SC; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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LA; OK; TX; South America (Argentina, Brazil, Paraguay, Uruguay) |
Discussion | The combination of spreading floral scales and scabrid culms separates this rather uncommon species from any others occurring in its range. The scarcity of Cyperus cephalanthus in the United States and its disjunct distribution suggest it might be naturalized rather than native. It was collected early in the nineteenth century in Louisiana, and the amphitropical distribution is not without parallel in the genus (cf. 12. C. eragrostis). Cyperus cephalanthus has been treated as a variety of the widespread, polymorphic South American C. laetus (G. Kükenthal 1935–1936). Recognition of C. cephalanthus as a species follows recent American floristic practice. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 23, p. 151. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. autumnalis, C. haspan subsp. juncoides, C. haspan var. americanus, C. juncoides | C. laetus var. cephalanthus |
Name authority | Linnaeus: Sp. Pl. 1: 45. (1753) | Torrey & Hooker: Ann. Lyceum Nat. Hist. New York. 3: 431. (1836) |
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