FNA: Cyperus grayoides vs. Cyperus sphaerolepis

	Cyperus grayoides	Cyperus sphaerolepis
	Illinois flatsedge, Mohlenbrock's sedge, umbrella sedge	Rusby's flatsedge, Rusby's sedge
Habit	Herbs, perennial, cespitose, rhizomatous, tuberlike.	Herbs, perennial, rhizomatous.
Culms	trigonous, 5–35 cm × 0.7–1.2 mm, glabrous.	basally cormlike, trigonous, (8–)15–40(–60) cm × (0.4–)0.8–1.5(–2) mm, glabrous or sparsely to densely scabridulous to minutely scabrid on angles or usually on surface, just proximal to bracts or over distal few centimeters.
Leaves	V-shaped, 5–25 cm × 1.5–3.5 mm, slightly scabridulous on abaxial margins, midrib.	(1–)3–4, V-shaped, (7–)15–25(–40) cm × 1–3(–5) mm, sparsely minutely scabridulous on margins and keel in distal 1/3, otherwise glabrous.
Inflorescences	spikes loosely ± globose, 15–25 mm wide; rays 1–10 cm; rachis 1–4 mm; bracts 3–7, ascending, V-shaped, 3–20 cm × 1–3 mm; rachilla deciduous, wingless.	spikes 1–3, sessile (occasionally 1 spike on peduncle 5–20 cm, produced from axil of distalmost leaf), loosely cylindric to oblong-ovoid, (9–)15–30(–45) × (3–)8–15(–18) mm; bracts 2–3(–7), longest erect, or very nearly so, others spreading, (1–)3–20(–26) cm × (0.5–)1–3(–5) mm; rays (1–)2–6(–11), (1–)2–7(–12) cm, in some depauperate plants rays less than 1 cm; rachilla ± deciduous, wingless.
Spikelets	3–24, compressed, oblong-lanceoloid, 4–12 × 2.5–3.2 mm; floral scales deciduous, 3–9, pale reddish brown, ovate, laterally 3–5-ribbed, 2.5–3 × 2–2.4 mm, apex acute to obtuse.	(4–)8–25(–35), oblong-ellipsoid to linear, compressed-quadrangular, (2–)7–11(–15) × (2–)2.4–3.2(–3.5) mm; floral scales (1–)4–8(–14), laterally dull whitish, stramineous, or light brown, or reddish brown to deep red, often red-dotted especially near keel, medially greenish, broadly ovate-deltate to ± orbiculate, laterally strongly 3(–4)-ribbed, medially weakly 1–3-ribbed, (1.6–)1.8–2.4 × (1.8–)2.2–2.8 mm, margins clear, erose especially apically, apex obtuse, tipped with straight mucro 0.1–0.2(–0.3) mm.
Flowers	anthers 1–1.2 mm; styles 0.4–0.6 mm; stigmas 1–2 mm.	anthers 0.4–0.6 mm; styles 0.4(–0.8) mm; stigmas 0.6–1 mm.
Achenes	dark brown, ± stipitate, ellipsoid, 2–2.4 × 0.9–1.2 mm, apex obtuse, apiculate, surfaces glabrous, puncticulate, or papillose.	finely reticulate with isodiametric cells, this layer gradually wearing away to reveal inner brown to reddish brown layer, stipitate, ovoid to obovoid, 1.4–1.6(–1.9) × 1–1.3(–1.4) mm, base cuneate to ± attenuate, stipe 0.1–0.2 × 0.2 mm, apex obtuse, slightly apiculate, inner layer papillose.
2n	= 166.

	Cyperus grayoides	Cyperus sphaerolepis
Phenology	Fruiting summer.	Fruiting summer.
Habitat	Sand prairies, waste places, fallow fields	Clearings in montane forests
Elevation	100–300 m (300–1000 ft)	1000–2500 m (3300–8200 ft)
Distribution	AR; IL; LA; MO; TX [WildflowerSearch map] [BONAP county map]	AZ; NM; TX; Mexico [BONAP county map]
Discussion	Of conservation concern. Cyperus grayoides is in the Center for Plant Conservation’s National Collection of Endangered Plant’s. See frontispiece. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Cyperus sphaerolepis has long been known by the synonym C. rusbyi and has been treated as a variety of C. fendlerianus. Specific status for C. sphaerolepis is supported by B. G. Marcks (1972) and G. C. Tucker (1994). (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23.	FNA vol. 23, p. 177.
Parent taxa	Cyperaceae > Cyperus > subg. Cyperus	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms		C. fendlerianus var. debilis, C. fendlerianus var. leucolepis, C. leucolepis, C. rusbyi, C. schweinitzii var. debilis
Name authority	Mohlenbrock: Brittonia 11: 255, fig. 1. (1959)	Boeckeler: Linnaea 35: 609. (1868)
Web links	Local Web sites: IL Wildflowers, LA Plants, MO Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus sphaerolepis

Illinois flatsedge, Mohlenbrock's sedge, umbrella sedge

Rusby's flatsedge, Rusby's sedge

Habit

Herbs, perennial, cespitose, rhizomatous, tuberlike.

Herbs, perennial, rhizomatous.

Culms

trigonous, 5–35 cm × 0.7–1.2 mm, glabrous.

basally cormlike, trigonous, (8–)15–40(–60) cm × (0.4–)0.8–1.5(–2) mm, glabrous or sparsely to densely scabridulous to minutely scabrid on angles or usually on surface, just proximal to bracts or over distal few centimeters.

Leaves

V-shaped, 5–25 cm × 1.5–3.5 mm, slightly scabridulous on abaxial margins, midrib.

(1–)3–4, V-shaped, (7–)15–25(–40) cm × 1–3(–5) mm, sparsely minutely scabridulous on margins and keel in distal 1/3, otherwise glabrous.

Inflorescences

spikes loosely ± globose, 15–25 mm wide;

rays 1–10 cm;

rachis 1–4 mm;

bracts 3–7, ascending, V-shaped, 3–20 cm × 1–3 mm;

rachilla deciduous, wingless.

spikes 1–3, sessile (occasionally 1 spike on peduncle 5–20 cm, produced from axil of distalmost leaf), loosely cylindric to oblong-ovoid, (9–)15–30(–45) × (3–)8–15(–18) mm;

bracts 2–3(–7), longest erect, or very nearly so, others spreading, (1–)3–20(–26) cm × (0.5–)1–3(–5) mm;

rays (1–)2–6(–11), (1–)2–7(–12) cm, in some depauperate plants rays less than 1 cm;

rachilla ± deciduous, wingless.

Spikelets

3–24, compressed, oblong-lanceoloid, 4–12 × 2.5–3.2 mm;

floral scales deciduous, 3–9, pale reddish brown, ovate, laterally 3–5-ribbed, 2.5–3 × 2–2.4 mm, apex acute to obtuse.

(4–)8–25(–35), oblong-ellipsoid to linear, compressed-quadrangular, (2–)7–11(–15) × (2–)2.4–3.2(–3.5) mm;

floral scales (1–)4–8(–14), laterally dull whitish, stramineous, or light brown, or reddish brown to deep red, often red-dotted especially near keel, medially greenish, broadly ovate-deltate to ± orbiculate, laterally strongly 3(–4)-ribbed, medially weakly 1–3-ribbed, (1.6–)1.8–2.4 × (1.8–)2.2–2.8 mm, margins clear, erose especially apically, apex obtuse, tipped with straight mucro 0.1–0.2(–0.3) mm.

Flowers

anthers 1–1.2 mm;

styles 0.4–0.6 mm;

stigmas 1–2 mm.

anthers 0.4–0.6 mm;

styles 0.4(–0.8) mm;

stigmas 0.6–1 mm.

Achenes

dark brown, ± stipitate, ellipsoid, 2–2.4 × 0.9–1.2 mm, apex obtuse, apiculate, surfaces glabrous, puncticulate, or papillose.

finely reticulate with isodiametric cells, this layer gradually wearing away to reveal inner brown to reddish brown layer, stipitate, ovoid to obovoid, 1.4–1.6(–1.9) × 1–1.3(–1.4) mm, base cuneate to ± attenuate, stipe 0.1–0.2 × 0.2 mm, apex obtuse, slightly apiculate, inner layer papillose.

= 166.

Cyperus grayoides

Cyperus sphaerolepis

Phenology

Fruiting summer.

Habitat

Sand prairies, waste places, fallow fields

Clearings in montane forests

Elevation

100–300 m (300–1000 ft)

1000–2500 m (3300–8200 ft)

Distribution

AR; IL; LA; MO; TX

[WildflowerSearch map]

[BONAP county map]

AZ; NM; TX; Mexico

[BONAP county map]

Discussion

Of conservation concern.

Cyperus grayoides is in the Center for Plant Conservation’s National Collection of Endangered Plant’s. See frontispiece.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus sphaerolepis has long been known by the synonym C. rusbyi and has been treated as a variety of C. fendlerianus. Specific status for C. sphaerolepis is supported by B. G. Marcks (1972) and G. C. Tucker (1994).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23.

FNA vol. 23, p. 177.

Parent taxa

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

C. fendlerianus var. debilis, C. fendlerianus var. leucolepis, C. leucolepis, C. rusbyi, C. schweinitzii var. debilis

Name authority

Mohlenbrock: Brittonia 11: 255, fig. 1. (1959)

Boeckeler: Linnaea 35: 609. (1868)

Web links

Local Web sites: IL Wildflowers, LA Plants, MO Plants
WildflowerSearch
iNaturalist (observations)
USDA Plants Database
LBJ Wildflower Center
SEINet
Plants of the World Online
Encyclopedia of Life
Wikipedia
Google Image Search

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Cyperus grayoides

Cyperus sphaerolepis

Cyperus grayoides

Cyperus sphaerolepis