FNA: Cyperus giganteus vs. Cyperus fendlerianus

	Cyperus giganteus	Cyperus fendlerianus
	giant flatsedge, Mexican papyrus	Fendler flatsedge, Fendler's flatsedge, Fendler's nutgrass, Fendler's sedge
Habit	Herbs, perennial, rhizomatous, stout.	Herbs, perennial, rhizomatous.
Culms	roundly trigonous, 200–350 cm × 15–50 mm, glabrous.	trigonous, (7–)20–70(–85) cm × (0.5–)1–1.8(–2.6) mm, glabrous or sparsely to densely scabridulous on surfaces or on angles immediately proximal to inflorescence in distal 3–6 cm.
Leaves	bladeless.	2–3, flat to V-shaped, (12–)20–40(–55) cm × (2–)3–5(–7) mm.
Inflorescences	spikes loosely cylindric, 15–40 × 8–16 mm; rays 5–8, ascending to erect, stiff,20–30 cm; 2d order rays 5–15 cm; bracts 10–12, ascending at 30–60°, flat, (8–) 20–45 cm × 4–12 mm; 2d order bracts (5–)15–25 cm × 1.5–5 mm; rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide.	spikes (1–)3–4(–6), largest erect, others at base of central spike, erect spike sessile, loosely to densely cylindric to ovoid-cylindric, (12–)18–30 × (6–)12–20 mm, remaining spikes 1/2 as large; rays absent; bracts (2–)3–6(–10), ± horizontal to reflexed parallel to culm, (2–)6–16(–25) cm × (1.5–)2–4(–7) mm; rachilla ± deciduous, wingless.
Spikelets	(15–)50–80, slightly compressed, linear, ± quadrangular, 5–12 × 0.6–1.4 mm; floral scales 8–20, appressed, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse.	(8–)15–30, spreading to ascending-appressed, oblong-lanceoloid, quadrangular, slightly compressed, 5–8(–10) × (2–)2.4–3.4(–4.2) mm; floral scales (2–)4–6(–8), yellowish, reddish, or greenish brown, medially greenish, straight to excurved, laterally (2–)3–4(–5)-ribbed, medially (1–)3–(–5)-ribbed, ovate-orbiculate, 2.4–2.8(–3.4) × 2.4–2.8(–3.4) mm, apex obtuse, mucronate to cuspidate, stout, (0.1–)0.3–0.6 mm, medially glabrous.
Flowers	anthers 0.7–1 mm (connective prolonged beyond anther as red subulate appendage 0.2–0.5 mm, its apex sometimes setose); styles 0.2–0.5 mm; stigmas (0.8–)1.2–1.8 mm.	anthers 0.4–0.8 mm; styles 0.7–1.3 mm; stigmas 0.7–1.2 mm.
Achenes	pale brown, sessile, oblong, 0.9–1 × 0.4–0.5 mm, apex scarcely apiculate, surfaces puncticulate.	finely reticulate, brown to reddish brown, ± stipelike, obovoid, 1.6–1.9 × 1.1–1.4(–1.6) mm, base attenuate to cuneate, 0.1–0.2 × 0.2 mm, apex obtuse, apiculate, surfaces glabrous.

	Cyperus giganteus	Cyperus fendlerianus
Phenology	Fruiting summer.	Fruiting summer.
Habitat	Stream banks, marshes	Clearings in montane forests
Elevation	0–10 m (0–0 ft)	500–1800 m (1600–5900 ft)
Distribution	LA; TX; Mexico; Central America; South America; West Indies [Introduced in North America] [BONAP county map]	AZ; CO; NM; TX; WY; Mexico [WildflowerSearch map] [BONAP county map]
Discussion	Cyperus giganteus was erroneously reported from Florida, a misidentification of C. papyrus (R. R. Haynes and A. Lasseigne 1969). Recently, some nurseries specializing in water plants have begun to offer C. giganteus under the name “Mexican papyrus.” Escaped cultivated plants may be expected in southeastern and southwestern United States. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Hybridization between Cyperus fendlerianus and C. sphaerolepis occurs occasionally in the southwestern United States and northern Mexico (B. G. Marcks 1972). The unnamed hybrid appears to be fully fertile and can be recognized by morphologic intermediacy. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 23, p. 173.	FNA vol. 23, p. 177.
Parent taxa	Cyperaceae > Cyperus > subg. Cyperus	Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens	C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms		Chlorocyperus fendlerianus
Name authority	Vahl: Enum. Pl. 2: 364. (1805)	Boeckeler: Linnaea 35: 520. (1868)
Web links	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cyperus fendlerianus

giant flatsedge, Mexican papyrus

Fendler flatsedge, Fendler's flatsedge, Fendler's nutgrass, Fendler's sedge

Habit

Herbs, perennial, rhizomatous, stout.

Herbs, perennial, rhizomatous.

Culms

roundly trigonous, 200–350 cm × 15–50 mm, glabrous.

trigonous, (7–)20–70(–85) cm × (0.5–)1–1.8(–2.6) mm, glabrous or sparsely to densely scabridulous on surfaces or on angles immediately proximal to inflorescence in distal 3–6 cm.

Leaves

bladeless.

2–3, flat to V-shaped, (12–)20–40(–55) cm × (2–)3–5(–7) mm.

Inflorescences

spikes loosely cylindric, 15–40 × 8–16 mm;

rays 5–8, ascending to erect, stiff,20–30 cm; 2d order rays 5–15 cm;

bracts 10–12, ascending at 30–60°, flat, (8–) 20–45 cm × 4–12 mm; 2d order bracts (5–)15–25 cm × 1.5–5 mm;

rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide.

spikes (1–)3–4(–6), largest erect, others at base of central spike, erect spike sessile, loosely to densely cylindric to ovoid-cylindric, (12–)18–30 × (6–)12–20 mm, remaining spikes 1/2 as large;

rays absent;

bracts (2–)3–6(–10), ± horizontal to reflexed parallel to culm, (2–)6–16(–25) cm × (1.5–)2–4(–7) mm;

rachilla ± deciduous, wingless.

Spikelets

(15–)50–80, slightly compressed, linear, ± quadrangular, 5–12 × 0.6–1.4 mm;

floral scales 8–20, appressed, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse.

(8–)15–30, spreading to ascending-appressed, oblong-lanceoloid, quadrangular, slightly compressed, 5–8(–10) × (2–)2.4–3.4(–4.2) mm;

floral scales (2–)4–6(–8), yellowish, reddish, or greenish brown, medially greenish, straight to excurved, laterally (2–)3–4(–5)-ribbed, medially (1–)3–(–5)-ribbed, ovate-orbiculate, 2.4–2.8(–3.4) × 2.4–2.8(–3.4) mm, apex obtuse, mucronate to cuspidate, stout, (0.1–)0.3–0.6 mm, medially glabrous.

Flowers

anthers 0.7–1 mm (connective prolonged beyond anther as red subulate appendage 0.2–0.5 mm, its apex sometimes setose);

styles 0.2–0.5 mm;

stigmas (0.8–)1.2–1.8 mm.

anthers 0.4–0.8 mm;

styles 0.7–1.3 mm;

stigmas 0.7–1.2 mm.

Achenes

pale brown, sessile, oblong, 0.9–1 × 0.4–0.5 mm, apex scarcely apiculate, surfaces puncticulate.

finely reticulate, brown to reddish brown, ± stipelike, obovoid, 1.6–1.9 × 1.1–1.4(–1.6) mm, base attenuate to cuneate, 0.1–0.2 × 0.2 mm, apex obtuse, apiculate, surfaces glabrous.

Cyperus giganteus

Cyperus fendlerianus

Phenology

Fruiting summer.

Habitat

Stream banks, marshes

Clearings in montane forests

Elevation

0–10 m (0–0 ft)

500–1800 m (1600–5900 ft)

Distribution

LA; TX; Mexico; Central America; South America; West Indies [Introduced in North America]

[BONAP county map]

AZ; CO; NM; TX; WY; Mexico

[WildflowerSearch map]

[BONAP county map]

Discussion

Cyperus giganteus was erroneously reported from Florida, a misidentification of C. papyrus (R. R. Haynes and A. Lasseigne 1969).

Recently, some nurseries specializing in water plants have begun to offer C. giganteus under the name “Mexican papyrus.” Escaped cultivated plants may be expected in southeastern and southwestern United States.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Hybridization between Cyperus fendlerianus and C. sphaerolepis occurs occasionally in the southwestern United States and northern Mexico (B. G. Marcks 1972). The unnamed hybrid appears to be fully fertile and can be recognized by morphologic intermediacy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 23, p. 173.

FNA vol. 23, p. 177.

Parent taxa

Cyperaceae > Cyperus > subg. Cyperus

Sibling taxa

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens

Synonyms

Chlorocyperus fendlerianus

Name authority

Vahl: Enum. Pl. 2: 364. (1805)

Boeckeler: Linnaea 35: 520. (1868)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Cyperus giganteus

Cyperus fendlerianus

Cyperus giganteus

Cyperus fendlerianus