The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

woodrush flatsedge

awn cyperus, awn flatsedge, bearded flatsedge

Habit Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. Herbs, annual, cespitose, with fibrous roots.
Culms

1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm.

1–20, trigonous, 2–16 cm × 0.5–0.8 mm, glabrous.

Leaves

(1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm.

1–3, flat to V-shaped, (1–)5–10(–15) × 0.5–2.5 mm.

Inflorescences

heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.;

rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm;

bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm.

spike 1, loosely to densely ovoid to oblong, 6–20(–40) × 9–15(–20) mm;

rays absent or 1–3(–6), 4–40 mm;

bracts (1–)2–4, longest ± erect, V-shaped, 1–15 cm × 0.5–3 mm;

rachilla ± deciduous, wingless.

Spikelets

(1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm;

floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous.

(2–)6–20(–40), greenish to reddish brown, somewhat compressed, ovoid-lanceoloid to oblong, 2.5–10(–20) × 1.3–2.2 mm (excluding awns);

floral scales deciduous, (4–)10–20(–34), greenish to stramineous or brownish red laterally, greenish medially, (5–)7–9(–11)-ribbed nearly to margins, oblong-lanceolate, (1.2–)1.3–1.8(–2.2) × (0.5–)0.7–0.8(–1) mm, apex cuspidate, excurved awn additional 0.5–1(–1.3) mm.

Flowers

stamen 1;

anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm;

styles 0.2–0.4 mm;

stigmas 0.4–0.6 mm.

stamen 1, filaments 1.5 mm;

anthers oblong, 0.2–0.3 mm, connective apex reddish, minute;

styles 0.3–0.5 mm;

stigmas 0.4–0.7 mm.

Achenes

brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate.

light brown to nearly black, stipitate, obovoid (occasionally linear-spatulate or linear oblong, infrequently constricted in middle), 0.7–0.8(–1.1) × (0.2–)0.3–0.4(–0.5) mm, stipe 0.05–0.1 × 0.1 mm, apex truncate, apiculate, surfaces papillose.

Cyperus entrerianus

Cyperus squarrosus

Phenology Fruiting summer. Fruiting summer.
Habitat Roadside ditches, marshes Moist, disturbed soils, gravelly roadsides, flood plains, edges of puddles, muddy places
Elevation 0–100 m (0–300 ft) 0–2500 m (0–8200 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; ON; QC; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America.

Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus squarrosus can be recognized by its small size and annual habit combined with its oblong-lanceolate floral scales bearing five to eleven conspicuous ribs and excurved awns. Some collections have been misidentified as C. acuminatus, an annual species of subg. Pycnostachys that has ovate-lanceolate, three-ribbed floral scales and digitately clustered spikelets.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 165.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. luzulae var. entrerianus Chlorocyperus inflexus, C. aristatus, C. aristatus var. inflexus, C. aristatus var. runyonii, C. inflexus, Dichostylis aristata, Mariscus squarrosus
Name authority Boeckeler: Flora 61: 139. (1878) Linnaeus: Cent. Pl. II, 6. (1756)
Web links