Cyperus entrerianus |
Cyperus refractus |
|
---|---|---|
woodrush flatsedge |
reflexed flatsedge |
|
Habit | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. | Herbs, perennial, cespitose, with inconspicuous, short rhizomes scarcely visible between culm bases. |
Culms | 1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
basally cormlike, trigonous, 30–100 cm × 2–4 mm, glabrous. |
Leaves | (1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
V-shaped to flat, 20–70 cm × 3–8 mm, adaxial surface, margins minutely scabridulous. |
Inflorescences | heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
spikes loose, broadly ellipsoid to ovoid, 4–7 × 2–6 cm; rays 6–13(–16), 4–25 cm, glabrous; rachis 1.5–3 cm; bracts 3–6, horizontal to ascending at 30°, flat, 5–35 cm × 2.5–6 mm; rachilla persistent, wings 0.4–0.5 mm wide. |
Spikelets | (1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
(13–)20–60(–74), linear, ± terete, (11–)13–25(–28) × 1.2–1.5 mm; distal spikelets spreading or ascending; floral scales persistent, (2–)3–6, stramineous, red-purple striate, laterally 3–5-ribbed, ovate-lanceolate, 4.5–5.3 × 1.2–1.5 mm. |
Flowers | stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
anthers 1–1.5 mm; styles 1.5 mm; stigmas 1.5 mm. |
Achenes | brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
brown to purplish brown, oblong-linear (5–6 times long as wide), 2.6–3 × 0.5–0.6 mm, base cuneate, apex obtuse, surfaces puncticulate. |
Cyperus entrerianus |
Cyperus refractus |
|
Phenology | Fruiting summer. | Fruiting summer (Jul–Aug). |
Habitat | Roadside ditches, marshes | Sandy riverbanks, gravelly roadsides, pastures, disturbed places |
Elevation | 0–100 m (0–300 ft) | 20–300 m (100–1000 ft) |
Distribution |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
|
AL; AR; DC; GA; MD; MO; NC; PA; SC; TN; VA; WV |
Discussion | Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus refractus is often mistaken for C. strigosus, which has shorter anthers (0.4–0.5 mm) and spreading, ± deciduous floral scales. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 182. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | |
Name authority | Boeckeler: Flora 61: 139. (1878) | Engelmann ex Boeckeler: Linnaea 36: 369. 1870, based on Cyperus refrofractus Engelmann ex Steudel, Syn. Pl. Glumac. 2: 47. 1855, not (Linnaeus) Torrey (1848) |
Web links |