Cyperus entrerianus |
Cyperus polystachyos |
|
---|---|---|
woodrush flatsedge |
many-spike flatsedge |
|
Habit | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. | Herbs, perennial (annual?), cespitose. |
Culms | 1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
trigonous or ± terete, (3–)20–45(–70) cm × (0.8–) 1.2–2 mm, glabrous. |
Leaves | (1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
(2–) 6–10(–16), flat to V-shaped, (2–) 10–30(–60) cm × 1–4 mm. |
Inflorescences | heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
spike 1, loosely ovoid to densely turbinate, 12–40 × 10–25 mm; rachis 3–10 mm; rays 1–8(–12), 0.5–3(–6) cm; 2d order rays absent; if rays absent, inflorescence single dense cluster of spikelets, 12–40 mm diam.; bracts (3–)4–6(–7), horizontal to ascending at 45°, flanged V-shaped, (2–)5–15(–23) cm × 1–4 mm. |
Spikelets | (1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
(5–)15–30(–40), spreading to appressed-ascending, linear-lanceoloid to linear, compressed, 8–18(–40) × (1–)1.3–1.6(–2) mm; floral scales (8–)12–25(–66), laterally stramineous to light brown, medially green, laterally ribless, medially 3–5-ribbed, oblong, (1.5–)1.8–2.4 × (1–)1.2–1.4 mm, apex acute to ± acute, mucronate. |
Flowers | stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
stamens 2; anthers (0.4–)0.5–0.8 mm, connective not prolonged; styles 0.6–1(–1.4) mm; stigmas (1–)1.4–2 mm. |
Achenes | brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
light to dark brown, oblong, (0.8–)0.9–1.1(–1.2) × 0.4–0.5(–0.6) mm, base cuneate to ± stipelike, apex truncate, apiculate, surfaces punctate to minutely punctate with raised cells. |
Cyperus entrerianus |
Cyperus polystachyos |
|
Phenology | Fruiting summer. | Fruiting summer–early fall. |
Habitat | Roadside ditches, marshes | Shores, ditches, swales between dunes |
Elevation | 0–100 m (0–300 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
|
AL; DC; DE; FL; GA; KY; LA; MA; MD; ME; MS; NC; NJ; NY; OK; PA; SC; TX; VA; WV; Mexico; Central America; South America; Asia; Africa
|
Discussion | Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus polystachyos is polymorphic. Adequate understanding of the infraspecific variations of C. polystachyos requires a worldwide examination, which is beyond the scope of this study. The segregates, C. fugax and C. filicinus, are recognized here; they have been usually accepted by recent workers (e.g., M. L. Fernald 1950; H. A. Gleason and A. Cronquist 1991). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 160. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | C. brizaeus, C. polystachyos var. leptostachyus, Pycreus polystachyos |
Name authority | Boeckeler: Flora 61: 139. (1878) | Rottbøll: Descr. Pl. Rar., 21. (1772) |
Web links |