Cyperus entrerianus
Cyperus fugax
woodrush flatsedge
withering flatsedge
1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm.
(1–)10–30(–60) together, roundly trigonous proximally, trigonous distally, (0.5–)3–6(–12) cm × (0.2–)0.4–0.8 mm, glabrous.
(1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm.
V-shaped, (1–)3–5(–10) cm × (0.5–)1–2 mm.
heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.;
rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm;
bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm.
spikes loosely ovoid, 8–15(–25) mm wide;
rachis 1–3 mm;
rays (1–)2–4(–6), (0.5–)1–3(–6) cm; 2d order rays absent;
bracts (1–)2–4(–5), ascending at 30–45°, V-shaped, (1–)2.5–6(–11) cm × 0.5–1.8 mm.
(1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm;
floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous.
(2–)4–6(–10), linear-lanceoloid, strongly flattened, (5–)8–12(–18) × 1.2–1.5 (1.8) mm;
floral scales (5–)10–16(–32), laterally clear, stramineous or light reddish brown, medially green, laterally ribless, medially 3(–5)-ribbed, ovate, (1.2–)1.3–1.6(–2) × 1–1.2(–1.4) mm, apex obtuse, mucronate to minutely mucronulate.
stamen 1;
anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm;
styles 0.2–0.4 mm;
stigmas 0.4–0.6 mm.
stamens 2;
anthers 0.1–0.2 mm;
styles 0.1–0.3(–0.4) mm;
stigmas 0.3–0.4 mm.
brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate.
light to dark brown, glossy or iridescent, stipitate, oblong-obovoid, 0.9–1.1 × 0.4–0.5 mm, stipe 0.1 mm, apex truncate, apiculate, surfaces punctate.
Cyperus entrerianus
Cyperus fugax
Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America.
Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Cyperus fugax is known from Louisiana from an early nineteeth century collection from “western Louisiana” (Hale s.n., TCD).
Cyperus fugax has been treated as a form of C. polystachyos (G. Kükenthal 1935–1936; A. B. Ayers 1946); nevertheless, specific status is appropriate. The most conspicuous differences are the annual habit and small size (less than 12 cm) of C. fugax; C. polystachyos is a perennial of moderate size (mostly taller than 20 cm). Other differences are summarized below.
Cyperus polystachyos is characterized as a shortly rhizomatous perennial; floral scales oblong, 1.8–2.4 mm, closely imbricate; styles 0.6–1 mm; stigmas 1.4–2 mm; anthers 0.6–0.8 mm; achenes substipitate to cuneate, slightly compressed laterally.
Cyperus fugax is characterized as a nonrhizomatous annual; floral scales ovate, 1.3–1.6 mm, loosely imbricate; styles 0.1–0.2 mm; stigmas 0.3–0.4 mm; filaments 1.2–1.4 mm; anthers 0.1–0.2 mm; achenes stipitate, strongly compressed laterally.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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