Cyperus entrerianus |
Cyperus flavescens |
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woodrush flatsedge |
pale flatsedge, yellow flatsedge |
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Habit | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. | Herbs, annual, cespitose. |
Culms | 1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
trigonous, (2–)4–30 cm × 0.4–2 mm, glabrous. |
Leaves | (1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
1–5, (blades often absent, base of culm with 1 reddish sheath bearing minute blade tooth 1–2 mm), (3–)10–18 cm × (0.5–)1.5–2(–2.6) mm. |
Inflorescences | heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
spikes 1–3, ovoid or ± digitate, 10–30 × 8–26 mm; rays 1–4(–6), 0.5–3(–9) cm; bracts 2–3, approximately horizontal, 1–12 cm × 0.5–2.5 mm. |
Spikelets | (1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
1–6, compressed, oblong-lanceoloid, 5–15 × (2–)2.3–2.8 mm; floral scales (4–)8–24, closely imbricate, laterally yellow to yellowish brown, margins light brown to clear, medially green, laterally ribless, medially 2–3-ribbed, 2-keeled basally, ovate, (1.5–)1.8–2.2 × 1.8 mm, apex obtuse. |
Flowers | stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
stamens 3; anthers 0.4 mm, connectives not prolonged; styles 0.5–1 mm; stigmas 0.5–0.8 mm. |
Achenes | brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
jet black to reddish brown, slightly stipitate, obovoid, 1–1.2 × 0.8 mm, apex apiculate, surface with network of rectangular longitudinally elongate cells and transverse undulations. |
Cyperus entrerianus |
Cyperus flavescens |
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Phenology | Fruiting summer. | Fruiting summer–early fall. |
Habitat | Roadside ditches, marshes | Damp, often disturbed soils |
Elevation | 0–100 m (0–300 ft) | 0–800 m (0–2600 ft) |
Distribution |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
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AL; AR; CA; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV; ON; South America; West Indies; Eurasia; Africa
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Discussion | Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
North American populations have been distinguished as Cyperus flavescens var. poiformis; their taxonomic separation was not supported by M. L. Corcoran’s (1941) careful study. Plants of Mexico and Central America consistently have purplish rather than yellowish floral scales and may be recognized as C. flavescens var. piceus (Liebmann) Fernald (G. C. Tucker 1983, 1994). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 162. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | C. flavescens var. poiformis, C. poiformis, Pycreus flavescens |
Name authority | Boeckeler: Flora 61: 139. (1878) | Linnaeus: Sp. Pl. 1: 46. (1753) |
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