Cyperus entrerianus |
Cyperus drummondii |
|
---|---|---|
woodrush flatsedge |
Drummond's sedge |
|
Habit | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. | Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. |
Culms | 1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
trigonous, 35–170 cm × 2–4.7 mm, scabrid on angles. |
Leaves | (1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
4–12, with readily visible cross ribs especially on abaxial surface, flat to V-shaped, 25–100 cm × 2.4–11 mm. |
Inflorescences | heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
heads digitate, globose, 1–1.7 cm diam.; rays 3–5, 1–7 cm; 2d order rays absent; bracts 3–5, horizontal to ascending at 30°, with prominent cross ribs, V-shaped, (4–)11–35(–48) cm × 1–6.7 mm (longest 1/2 as long as culm); 2d order bracts absent. |
Spikelets | (1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
10–40, oblong to linear-lanceoloid, 5–1.8 × 1.5–2.2 mm; floral scales 18–42, pale grayish green, turning brownish, proximally greenish or brownish, 2-keeled, weakly to distinctly 1-ribbed, proximal 1/2 2-ribbed, ovate, 1.4–1.6 × 0.9–1.4 mm, acute (sometimes mucronulate), apically glabrous, occasionally scabridulous. |
Flowers | stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
stamens 1–2; anthers 0.8–1.2 mm; styles 0.8–1 mm; stigmas 0.6–0.8 mm. |
Achenes | brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
brown, slightly stipitate, oblong-ellipsoid (2.4–3.9 times longer than wide), 1–1.2 × 0.3–0.5 mm, apical beak 0.1–0.5 mm, surfaces glabrous. |
Cyperus entrerianus |
Cyperus drummondii |
|
Phenology | Fruiting summer. | Fruiting late spring–summer. |
Habitat | Roadside ditches, marshes | Wet, relatively undisturbed habitats, especially flatwoods, ponds, seepage slopes, coastal prairies |
Elevation | 0–100 m (0–300 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
|
AL; FL; GA; LA; MS; SC; TX; Central America; South America; West Indies |
Discussion | Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus drummondii has been treated as a synonym (G. C. Tucker 1994), or a variety (G. Kükenthal 1935–1936), or a subspecies of C. virens (M. F. Denton 1978b). Recent quantitative and field studies (J. R. Carter et al. 1999) provided compelling evidence that specific status is appropriate. Compared to Cyperus virens, C. drummondii is a taller species of less disturbed habitats and has narrower spikelets, ovate scales, and fewer inflorescence bracts. Cyperus virens and C. drummondii are readily distinguished from other Cyperus species in the United States by their sharply angled, scabrid culms and conspicuously septate leaf blades. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | C. robustus, C. virens subsp. drummondii, C. virens var. drummondii, C. virens var. robustus |
Name authority | Boeckeler: Flora 61: 139. (1878) | Torrey & Hooker: Ann. Lyceum Nat. Hist. New York 3: 437, errata. (1836) |
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