Cyperus entrerianus |
Cyperus distans |
|
---|---|---|
woodrush flatsedge |
Piedmont flatsedge |
|
Habit | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. | Herbs, perennial, rhizomatous. |
Culms | 1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
single or close together, trigonous, (10–)40–80(–120) cm × 1.2–2.5(–5) mm, glabrous. |
Leaves | (1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
2–5, pleated, (10–)20–50(–90) cm × 2–5(–10) mm. |
Inflorescences | heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
spikes 1–4(–7), loose, broadly ovoid to ovoid-pyramidal, 25–35 × 25–35 mm; rays (4–)5–6(–8), (1–)3–8(–16) cm, glabrous; 2d order rays (0–)1–5, 5–15(–50) mm; bracts 4–7, ascending at 30–45°, pleated, (5–)20–40(–60) × (2–)3–8(–11) mm; 2d order bracts 1–4 per ray, 5–10(–30) × 0.5–1 mm, scabridulous; rachilla persistent, wingless (or very narrowly winged, not clasping achene). |
Spikelets | (1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
(10–)20–35(–45), seemingly flattened proximally from spreading floral scales, becoming terete apically, linear, (10–)13–18(–25) × 1.2–1.5(–1.6) mm; floral scales deciduous, (10–)12–20(–25), laterally light reddish brown to dark red, medially greenish, laterally ribless, medially 3–5-ribbed, oblong-ovate, 1.7–2(–2.4) × 0.9–1.2 mm, apex obtuse, barely reaching next scale, apex entire, emarginate, mucro at most 0.3 mm. |
Flowers | stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
anthers 0.4–0.5 mm; styles 0.3–0.6 mm; stigmas 1–1.8(–2) mm. |
Achenes | brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
dark brown, narrowly oblong, (1.3–)1.4–1.6(–1.7) × (0.3–)0.4–0.5 mm, base cuneate to ± stipelike, apex acute, not apiculate, surfaces papillose. |
Cyperus entrerianus |
Cyperus distans |
|
Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Roadside ditches, marshes | Marshes |
Elevation | 0–100 m (0–300 ft) | 0 m (0 ft) |
Distribution |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
|
FL; NC; Mexico; South America; West Indies (Dominican Republic south through the Lesser Antilles); Asia; Africa |
Discussion | Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 23. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | C. elatus |
Name authority | Boeckeler: Flora 61: 139. (1878) | Linnaeus f.: Suppl. Pl., 103. (1782) |
Web links |