Cyperus entrerianus |
Cyperus diandrus |
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woodrush flatsedge |
umbrella flat sedge |
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Habit | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. | Herbs, annual, cespitose, with fibrous roots. |
Culms | 1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
trigonous, 4–25(–35) cm, glabrous. |
Leaves | (1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
flat, 5–20 cm × 1.5–3 mm. |
Inflorescences | heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
heads ± digitate, 12–22 mm diam.; rays 3–6, 1–6 cm; bracts 1–4, ± horizontal, flat, 1.5–15 cm × 1.5–3 mm. |
Spikelets | (1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
6–12, oblong-lanceoloid, compressed, 5–10 × 2.8–3.2 mm; floral scales 8–28, closely imbricate, stramineous to light brown or clear (marginal band bright red, spikelet thus appearing dark margined), laterally ribless, oblong to ovate, 2.5–3 × 1.6–1.9 mm. |
Flowers | stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
stamens 2(–3); anthers 0.3 mm, connective apex reddish, subulate, 0.1–0.2 mm; styles 0.3–0.5 mm; stigmas 2.2–3.1 mm. |
Achenes | brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
brown, network of ridges forming isodiametric or square cells, sessile or stipitate, obovoid to ovoid, 1–1.2 × 0.5–0.6 mm, stipe, if present, 0.1 mm, apex obtuse, apiculate, surfaces minutely punctate. |
Cyperus entrerianus |
Cyperus diandrus |
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Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Roadside ditches, marshes | Emergent shorelines, in sandy, peaty, or slightly brackish areas (though seldom where disturbed) |
Elevation | 0–100 m (0–300 ft) | 0–600 m (0–2000 ft) |
Distribution |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
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CT; DE; IA; IL; IN; MA; MD; ME; MI; MN; MO; NE; NH; NJ; NY; PA; RI; VA; VT; NB; NS; ON; QC
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Discussion | Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 23. | FNA vol. 23, p. 161. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Pycreus |
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | Pycreus diander |
Name authority | Boeckeler: Flora 61: 139. (1878) | Torrey: Cat. Pl. New York, 90. (1819) |
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