Cyperus entrerianus |
Cyperus dentoniae |
|
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woodrush flatsedge |
Denton's flatsedge, hairy flatsedge |
|
Habit | Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. | Herbs, perennial, cespitose, shortly rhizomatous. |
Culms | 1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm. |
1–4, trigonous, (2–)20–50(–80) cm × (0.8–)1.5–2(–2.8) mm, basally glabrous or nearly so, apically hispidulous, on faces and angles distally, especially so immediately proximal to bracts. |
Leaves | (1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm. |
2–5(–7), V-shaped, (5–)15–30(–45) cm × (1.5–)2.5–5(–8) mm. |
Inflorescences | heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.; rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm; bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm. |
spike 1 (occasionally 1–2 smaller sessile spikes at base), densely cylindric, (7–)11–20(–25) × (5–) 7–10(–12) mm; rays 3–6(–10), (1–)3–6(–18) cm, rays and rachis hispidulous; bracts 4–8(–10), ascending at 30–60°, V-shaped, (2–)10–25(–45) cm × (1–)3–6(–8) mm; rachilla deciduous, wings persistent, 0.3–0.4 mm wide. |
Spikelets | (1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm; floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous. |
(20–)70–120(–140), oblong-ellipsoid to oblong-lanceoloid, ± terete to quadrangular, (2.2–)3.4–4.5(–5.6) × (0.8–)0.9–1.2(–1.3) mm; distal spikelet spreading or ascending; floral scales 1(–3), appressed, stramineous, red-spotted, strongly 3(–4)-ribbed on either side of green part, medially weakly 3-ribbed, ovate to elliptic ovate, (2.4–)2.8–3.3(–3.6) × 1.6–2 mm, apex acute, mucronulate from excurrent midrib. |
Flowers | stamen 1; anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm; styles 0.2–0.4 mm; stigmas 0.4–0.6 mm. |
anthers (0.3–)0.4–0.6 mm; styles 0.4–0.6(–0.8) mm; stigmas 1–1.4 mm. |
Achenes | brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate. |
brown, stipitate, ellipsoid, (1.6–)1.8–2 × (0.7–) 0.8–0.9 mm, stipe 0.1–0.2 × 0.2 mm, apex ± truncate, apiculate, surfaces puncticulate to glabrous. |
Cyperus entrerianus |
Cyperus dentoniae |
|
Phenology | Fruiting summer. | Fruiting mid summer–fall (Jul–Sep). |
Habitat | Roadside ditches, marshes | Various dry, open environments |
Elevation | 0–100 m (0–300 ft) | 500–1000 m (1600–3300 ft) |
Distribution |
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
|
AZ; Mexico; Central America; South America |
Discussion | Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America. Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus dentoniae was not treated in G. Kükenthal’s monograph (1935–1936), apparently because he did not see any materials of the species. The plant has been treated as “C. asper (Liebmann) O’Neill,” a name based on Mariscus asper Liebmann and a synonym of C. mutisii (A. B. Ayers 1946). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 187. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. luzulae var. entrerianus | Mariscus pubescens, C. flavomariscus var. peduncularis, C. flavus var. peduncularis |
Name authority | Boeckeler: Flora 61: 139. (1878) | G. C. Tucker: Syst. Bot. Monogr. 2: 56. (1983) |
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