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woodrush flatsedge

cyperus, flatsedge, galingale, nutsedge, umbrella-sedge

Habit Herbs, perennial, coarse; rhizomes (seldom collected) indurate, oblique, 5–12 mm wide, with fibrous brown floral scales, 1–5 cm. Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous.
Culms

1–3, trigonous to roundly trigonous, (30–)40–65(–95) cm × (1–)2–3 mm.

solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles.

Leaves

(1–)5–10, V-shaped, (10–)30–45(–70) cm × 3–7 mm.

usually basal;

ligules absent;

blades keeled abaxially, flat, V-, or inversely W-shaped in cross section.

Inflorescences

heads loosely to densely globose-ovoid, (6–)10–12(–20) mm diam.;

rays 6–10(–12), 1–8(–12) cm; 2d order rays 1–4, 5–15(–23) mm;

bracts (5–)6–8(–10), ascending at 45–60(–75)°, V-shaped, (5–)15–40(–55) cm × (1–)3–7 mm.

terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet;

spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts;

involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike.

Spikelets

(1–)30–50(–65), greenish white, linear to broadly ellipsoid, flattened, 4–6.5 × 1.8–3.2 mm;

floral scales (10–)16–20(–26), laterally clear, pale green, off-white, or light brown, medially green or light brown, laterally weakly 1-ribbed, medially 2-ribbed, basally 2-keeled, oblong-lanceolate, (1.2–)1.4–1.6(–1.8) × (0.7–)0.8–1 mm, apex acute, mucronate, distally glabrous or scabridulous.

scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous).

Flowers

stamen 1;

anthers ellipsoid, 0.6–0.7 mm, connective apex acute, 0.1–0.2 mm;

styles 0.2–0.4 mm;

stigmas 0.4–0.6 mm.

bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base deciduous or persistent;

stigmas 2–3.

Achenes

brown, stipitate, ellipsoid to narrowly ellipsoid, broadly rounded or contracted basally, gradually tapered toward apex, 0.9–1.1 × 0.3–0.4 mm, stipe 0.1 mm, apex acute, surfaces finely reticulate.

biconvex, flattened, or trigonous.

Cyperus entrerianus

Cyperus

Phenology Fruiting summer.
Habitat Roadside ditches, marshes
Elevation 0–100 m (0–300 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; TX; Mexico; South America; Central America (Nicaragua) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from USDA
Pantemperate and tropical
[BONAP county map]
Discussion

Cyperus entrerianus is a recent introduction in the southern United States (earliest collection from Pensacola County, Florida, Brinker 413, MO in 1941). It has been confused with C. luzulae. The species that Boeckeler had first described 50 years earlier was accepted by G. Kükenthal (1935–1936). Cyperus entrerianus was accepted as a variety of C. luzulae by A. B. Ayers (1946); M. F. Denton (1978b) considered C. entrerianus a synonym of C. luzulae, stating that the features of these two taxa merged so completely throughout their geographic ranges that C. entrerianus could not be given specific status. Cyperus luzulae in the strict sense does not extend northward to North America.

Morphologic differences between Cyperus entrerianus and C. luzulae are as follows (G. C. Tucker 1994). Cyperus entrerianus has culms 40–75 cm; inflorescence bracts ascending at 45–60°; primary rays ascending at 45–60(–75)°; 2d order rays present; heads of spikelets loosely globose-ovoid, light greenish white to golden brown; rachilla dark red with stramineous scale scars; and scale apices acute. Cyperus luzulae has culms 20–40 cm; inflorescence bracts approximately horizontal; primary rays ascending at 15–30°; 2d order rays absent; heads of spikelets densely oblong to pyramidal, bright to dull white; rachilla green throughout; and scale apices rounded.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 600 (96 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas 2 (3 in C. serotinus); achenes biconvex.
→ 2
1. Stigmas 3; achenes trigonous, plano-convex, or terete.
→ 3
2. Achenes laterally flattened, borne with edge toward rachilla.
subg. Pycreus
2. Achenes dorsiventrally flattened, borne with face toward rachilla.
subg. Juncellus
3. Spikelets borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads.
subg. Pycnostachys
3. Spikelets borne in spikes on conspicuous rachis.
→ 4
4. Rachilla continuous or articulate only at base.
subg. Cyperus
4. Rachilla articulate at base of each scale, mature spikelet disarticulating into segments consisting of scale, internode, and rachilla wings.
subg. Diclidium
Source FNA vol. 23. FNA vol. 23, p. 141. Authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Subordinate taxa
C. subg. Cyperus, C. subg. Diclidium, C. subg. Juncellus, C. subg. Pycnostachys, C. subg. Pycreus
Synonyms C. luzulae var. entrerianus
Name authority Boeckeler: Flora 61: 139. (1878) Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754)
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