Cyperus echinatus |
Cyperus retroflexus |
|
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globe flatsedge, teasel sedge |
one-flower flatsedge |
|
Habit | Herbs, perennial, single-stemmed to loosely cespitose. | Herbs, perennial, cespitose, rhizomatous. |
Culms | basally cormlike, trigonous, (15–)30–100 cm × 0.5–3.5 mm, glabrous. |
trigonous, basally tuberous-thickened, (5–)15–40(–80) cm × 1–2.5 mm, glabrous. |
Leaves | flat to V-shaped, 10–65 cm × 3–9 mm, adaxial surface, margins minutely scabridulous. |
flat to V-shaped, (5–)10–30(–60) cm × (0.5–)1–3(–4) mm. |
Inflorescences | spikes densely globose to globose-ovoid, 8–17 mm wide; rays 3–12, 2–12 cm, scaberous adaxially especially distally; rachis 4–8 mm; bracts (3–)4–7, ascending at 30(–45)°, flat, 5–35 cm × 2–9 mm; rachilla persistent, wings 0.5–0.7 mm wide. |
spike 1, loosely to densely ovoid to cylindric, 8–36 × 8–20 mm; rays 3–6(–9), (5–)10–32 cm; bracts 3–6, horizontal to ascending at 30°, V-shaped, (1–)3–15(–30) cm × 0.5–4.5 mm; rachilla deciduous, wings 0.5–0.7 mm wide. |
Spikelets | 50–100, oblong-lanceoloid, ± terete-quadrangular, (3.5–)4–7 × 1–1.4 mm; distal spikelet spreading or ascending; floral scales persistent, 3–5, appressed, stramineous to brownish, 4-ribbed laterally, oblong-elliptic, 3.5–4.5 × 1–1.8 mm, membranous, apex entire or emarginate with mucro to 0.3 mm. |
(5–)20–35(–65), linear to oblong (without narrowed basal portion), quadrangular; (2.2–)4–10(–18) × 0.7–1.1 mm; floral scales persistent, 1–5, appressed, pale greenish white laterally, stramineous with reddish dots, or deep red, greenish or light brown medially, 3–6-ribbed laterally, narrowly ovate to oblong-elliptic, 2.6–3.9 × 1.6–2 mm, apex rounded to ± acute, entire, mucronulate; distal scale with glabrous midrib and mucro 0.1–0.3(–0.5) mm; sterile terminal scale of spikelet only 0.4–0.6 mm wide, involute, frequently uncinate. |
Flowers | anthers 0.4–0.8 mm; styles 0.5–0.6 mm; stigmas 1 mm. |
anthers (0.3–)0.4–0.6 mm; styles 0.4–0.5 mm; stigmas 0.5–0.9 mm. |
Achenes | brown, ± stipitate, oblong, (1.5–)1.8–2.3 × 0.5–0.6(–0.7) mm (1/2 length of floral scales), apex obtuse, surfaces puncticulate. |
light brown to reddish brown, sessile or slightly stipitate, ellipsoid, (1.8–)2–2.4 × 0.6–0.8(–0.9) mm, apex slightly apiculate, surfaces puncticulate. |
Cyperus echinatus |
Cyperus retroflexus |
|
Phenology | Fruiting summer–early fall. | Fruiting mid summer–early fall (Jul–Sep). |
Habitat | Disturbed, sunny sites, in mesic places, well-drained soils | Damp, disturbed soils, croplands |
Elevation | 0–500 m (0–1600 ft) | 0–1000 m (0–3300 ft) |
Distribution |
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; West Indies
|
AL; AR; AZ; MO; MS; NM; OK; TX; Mexico
|
Discussion | The records for Rhode Island and Wisconsin are according to M. L. Horvat (1941); we have not seen specimens from those states. Cyperus echinatus is usually recognized by its tight, nearly spheric spikes; it may occasionally be hard to distinguish from C. croceus and C. retrorsus. Compared to C. retrorsus, C. echinatus has larger spikelets and longer floral scales, anthers, and achenes. In contrast to C. echinatus, C. croceus has looser spikes, shorter, broader, greenish or yellowish floral scales, shorter, more ovoid achenes, and shorter anthers. Furthermore, C. echinatus is predominantly an inland species of roadsides, pastures, and other disturbed ground; C. retrorsus is primarily a coastal species and occurs in drier, sandier sites. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus retroflexus is distinguished from other species of Cyperus with deciduous spikelets by the involute, sterile terminal scale of the spikelet. In plants with cuspidate scales, the terminal scale is usually somewhat recurved, giving the appearance of a hook at the end of the spikelet. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 182. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | Scirpus echinatus, C. ovularis, C. ovularis var. americanus, C. ovularis var. sphaericus, C. ovularis var. wolfii, C. wolfii, Kyllinga ovularis, Mariscus ovularis | C. uniflorus var. floribundus, C. uniflorus var. retroflexus, Mariscus uniflorus |
Name authority | (Linnaeus) Alph. Wood: Class-book Bot. ed. s.n.(b), 734. (1861) | Buckley: Proc. Acad. Nat. Sci. Philadelphia. 14: 9. (1862) |
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