Cyperus echinatus |
Cyperus reflexus |
|
---|---|---|
globe flatsedge, teasel sedge |
bentawn flatsedge |
|
Habit | Herbs, perennial, single-stemmed to loosely cespitose. | Herbs, perennial, cespitose; rhizomes elongate, scaly, 2–3 mm wide. |
Culms | basally cormlike, trigonous, (15–)30–100 cm × 0.5–3.5 mm, glabrous. |
roundly trigonous to terete, 30–80 cm × 1–1.6 mm, glabrous. |
Leaves | flat to V-shaped, 10–65 cm × 3–9 mm, adaxial surface, margins minutely scabridulous. |
1–6, 10–40 cm, glabrous or essentially so. |
Inflorescences | spikes densely globose to globose-ovoid, 8–17 mm wide; rays 3–12, 2–12 cm, scaberous adaxially especially distally; rachis 4–8 mm; bracts (3–)4–7, ascending at 30(–45)°, flat, 5–35 cm × 2–9 mm; rachilla persistent, wings 0.5–0.7 mm wide. |
heads ovoid to pyramidal, 9–15 mm diam.; rays 0 or 3–6, 0.5–4 cm, glabrous; bracts 3–4, longest erect, appearing as continuation of culm, others horizontal to ascending, 4–17 cm × 1.5–4 mm. |
Spikelets | 50–100, oblong-lanceoloid, ± terete-quadrangular, (3.5–)4–7 × 1–1.4 mm; distal spikelet spreading or ascending; floral scales persistent, 3–5, appressed, stramineous to brownish, 4-ribbed laterally, oblong-elliptic, 3.5–4.5 × 1–1.8 mm, membranous, apex entire or emarginate with mucro to 0.3 mm. |
25–60, reddish with contrasting yellowish or greenish edges, oblong-lanceoloid, strongly compressed, 5–6(–15) × 1.5–2.5 mm; floral scales 10–22, laterally reddish, medially yellowish green, 2-keeled, laterally strongly 1-ribbed on each side, proximally grooved, ovate-elliptic, 1.4–2.2 × 0.8–1.5 mm, apex acute, apically scabridulous (sometimes glabrous). |
Flowers | anthers 0.4–0.8 mm; styles 0.5–0.6 mm; stigmas 1 mm. |
stamen 1; anthers 0.6–1 mm; styles 0.3–0.5 mm; stigmas 0.3–0.6 mm. |
Achenes | brown, ± stipitate, oblong, (1.5–)1.8–2.3 × 0.5–0.6(–0.7) mm (1/2 length of floral scales), apex obtuse, surfaces puncticulate. |
reddish brown, stipitate, ovoid-ellipsoid, 0.8–0.9 × 0.3–0.4 mm, stipe 0.1–0.2 mm, apex acute to acuminate, surfaces reticulate or subtly rugulose. |
Cyperus echinatus |
Cyperus reflexus |
|
Phenology | Fruiting summer–early fall. | Fruiting late spring–fall (May–Oct). |
Habitat | Disturbed, sunny sites, in mesic places, well-drained soils | Muddy soils or shallow waters |
Elevation | 0–500 m (0–1600 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; West Indies
|
FL; LA; OK; TX; Mexico; Central America (Costa Rica) |
Discussion | The records for Rhode Island and Wisconsin are according to M. L. Horvat (1941); we have not seen specimens from those states. Cyperus echinatus is usually recognized by its tight, nearly spheric spikes; it may occasionally be hard to distinguish from C. croceus and C. retrorsus. Compared to C. retrorsus, C. echinatus has larger spikelets and longer floral scales, anthers, and achenes. In contrast to C. echinatus, C. croceus has looser spikes, shorter, broader, greenish or yellowish floral scales, shorter, more ovoid achenes, and shorter anthers. Furthermore, C. echinatus is predominantly an inland species of roadsides, pastures, and other disturbed ground; C. retrorsus is primarily a coastal species and occurs in drier, sandier sites. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus reflexus is recognized by its reddish scales with contrasting greenish or yellow keels and by the laterally 1ribbed scales. The lateral rib is most often situated where the reddish and yellowish parts of the scale meet. Two varieties of C. reflexus, var. reflexus and var. fraternus, have been recognized (M. F. Denton 1978b). Both varieties were noted as intermingled in the North American and South American segments of the widely disjunct range of C. reflexus. Such variation does not seem significant at the varietal level (see G. C. Tucker 1994; G. C. Tucker and R. McVaugh 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 182. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Pycnostachys |
Sibling taxa | ||
Synonyms | Scirpus echinatus, C. ovularis, C. ovularis var. americanus, C. ovularis var. sphaericus, C. ovularis var. wolfii, C. wolfii, Kyllinga ovularis, Mariscus ovularis | C. arenicola, C. fraternus, C. pseudovegetus var. arenicola, C. reflexus var. fraternus |
Name authority | (Linnaeus) Alph. Wood: Class-book Bot. ed. s.n.(b), 734. (1861) | Vahl: Enum. Pl. 2: 299. (1805) |
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