Cyperus echinatus |
Cyperus aggregatus |
|
---|---|---|
globe flatsedge, teasel sedge |
ballast sedge, inflated-scale flatsedge |
|
Habit | Herbs, perennial, single-stemmed to loosely cespitose. | Herbs, perennial, cespitose, rhizomatous. |
Culms | basally cormlike, trigonous, (15–)30–100 cm × 0.5–3.5 mm, glabrous. |
trigonous, 20–100 cm × 0.8–2.5 mm, glabrous. |
Leaves | flat to V-shaped, 10–65 cm × 3–9 mm, adaxial surface, margins minutely scabridulous. |
5–10, V-shaped to flanged V-shaped, 10–70(–90) cm × 2–7 mm, margins and midribs scabridulous or glabrous. |
Inflorescences | spikes densely globose to globose-ovoid, 8–17 mm wide; rays 3–12, 2–12 cm, scaberous adaxially especially distally; rachis 4–8 mm; bracts (3–)4–7, ascending at 30(–45)°, flat, 5–35 cm × 2–9 mm; rachilla persistent, wings 0.5–0.7 mm wide. |
spikes 1(–6), densely cylindric, 6–30 × 5–11 mm; rays 3–12, 0.4–5(–7) cm; rays and rachis glabrous; usually only 1–2 spikes of inflorescence on elongate rays, other spikes sessile or nearly so; bracts 4–7, horizontal to slightly ascending, 1–16 cm × 0.5–4 mm; rachilla deciduous, wings persistent, 0.5 mm wide. |
Spikelets | 50–100, oblong-lanceoloid, ± terete-quadrangular, (3.5–)4–7 × 1–1.4 mm; distal spikelet spreading or ascending; floral scales persistent, 3–5, appressed, stramineous to brownish, 4-ribbed laterally, oblong-elliptic, 3.5–4.5 × 1–1.8 mm, membranous, apex entire or emarginate with mucro to 0.3 mm. |
20–80, ellipsoid, roughly quadrangular, 3–5 × 1–1.4 mm; floral scales 1–2(–4), appressed, stramineous to golden brown, often red-speckled, medially greenish, 9-ribbed, elliptic to ovate, 2.4–3.4 × 1.8–2.6 mm, apex obtuse. |
Flowers | anthers 0.4–0.8 mm; styles 0.5–0.6 mm; stigmas 1 mm. |
anthers 0.4–0.6 mm; styles 0.7–1.1 mm; stigmas 1.4–2.1 mm. |
Achenes | brown, ± stipitate, oblong, (1.5–)1.8–2.3 × 0.5–0.6(–0.7) mm (1/2 length of floral scales), apex obtuse, surfaces puncticulate. |
dark brown to reddish brown, sessile, broadly ellipsoid, 1.8–2.1 × 0.8–1 mm, apex apiculate, surfaces glabrous to finely puncticulate. |
Cyperus echinatus |
Cyperus aggregatus |
|
Phenology | Fruiting summer–early fall. | Fruiting mid summer–fall (Jul–Oct). |
Habitat | Disturbed, sunny sites, in mesic places, well-drained soils | Dry roadsides, pastures, thickets |
Elevation | 0–500 m (0–1600 ft) | 0–500 m (0–1600 ft) |
Distribution |
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; West Indies
|
AZ; LA; MS; NJ; NM; OR; PA; TX; Mexico; Central America; South America [Introduced, Australia]
|
Discussion | The records for Rhode Island and Wisconsin are according to M. L. Horvat (1941); we have not seen specimens from those states. Cyperus echinatus is usually recognized by its tight, nearly spheric spikes; it may occasionally be hard to distinguish from C. croceus and C. retrorsus. Compared to C. retrorsus, C. echinatus has larger spikelets and longer floral scales, anthers, and achenes. In contrast to C. echinatus, C. croceus has looser spikes, shorter, broader, greenish or yellowish floral scales, shorter, more ovoid achenes, and shorter anthers. Furthermore, C. echinatus is predominantly an inland species of roadsides, pastures, and other disturbed ground; C. retrorsus is primarily a coastal species and occurs in drier, sandier sites. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The records of Cyperus aggregatus in New Jersey, Oregon, and Pennsylvania represent introductions from ballast. The closely related Mexican and Central American Cyperus regiomontanus Britton has been collected once as an adventive or waif in southern California. It is distinguished from C. aggregatus by its silvery green, subulate-lanceolate spikelets and achenes that are only 0.4–0.6 mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 182. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | Scirpus echinatus, C. ovularis, C. ovularis var. americanus, C. ovularis var. sphaericus, C. ovularis var. wolfii, C. wolfii, Kyllinga ovularis, Mariscus ovularis | Mariscus aggregatus, C. cayennensis, C. flavus, C. flavus var. aggregatus, C. flavus var. laevis, Kyllinga cayennensis, Kyllinga squarrosa, Mariscus flavus, Mariscus laevis |
Name authority | (Linnaeus) Alph. Wood: Class-book Bot. ed. s.n.(b), 734. (1861) | (Willdenow) Endlicher: Cat. Horti Vindob. 1: 93. (1842) |
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