Cyperus echinatus |
Cyperus |
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globe flatsedge, teasel sedge |
cyperus, flatsedge, galingale, nutsedge, umbrella-sedge |
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Habit | Herbs, perennial, single-stemmed to loosely cespitose. | Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous. | ||||||||||||||||
Culms | basally cormlike, trigonous, (15–)30–100 cm × 0.5–3.5 mm, glabrous. |
solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles. |
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Leaves | flat to V-shaped, 10–65 cm × 3–9 mm, adaxial surface, margins minutely scabridulous. |
usually basal; ligules absent; blades keeled abaxially, flat, V-, or inversely W-shaped in cross section. |
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Inflorescences | spikes densely globose to globose-ovoid, 8–17 mm wide; rays 3–12, 2–12 cm, scaberous adaxially especially distally; rachis 4–8 mm; bracts (3–)4–7, ascending at 30(–45)°, flat, 5–35 cm × 2–9 mm; rachilla persistent, wings 0.5–0.7 mm wide. |
terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet; spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts; involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike. |
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Spikelets | 50–100, oblong-lanceoloid, ± terete-quadrangular, (3.5–)4–7 × 1–1.4 mm; distal spikelet spreading or ascending; floral scales persistent, 3–5, appressed, stramineous to brownish, 4-ribbed laterally, oblong-elliptic, 3.5–4.5 × 1–1.8 mm, membranous, apex entire or emarginate with mucro to 0.3 mm. |
scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous). |
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Flowers | anthers 0.4–0.8 mm; styles 0.5–0.6 mm; stigmas 1 mm. |
bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings; perianth absent; stamens 1–3; styles linear, 2–3-fid, base deciduous or persistent; stigmas 2–3. |
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Achenes | brown, ± stipitate, oblong, (1.5–)1.8–2.3 × 0.5–0.6(–0.7) mm (1/2 length of floral scales), apex obtuse, surfaces puncticulate. |
biconvex, flattened, or trigonous. |
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Cyperus echinatus |
Cyperus |
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Phenology | Fruiting summer–early fall. | |||||||||||||||||
Habitat | Disturbed, sunny sites, in mesic places, well-drained soils | |||||||||||||||||
Elevation | 0–500 m (0–1600 ft) | |||||||||||||||||
Distribution |
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; West Indies
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Pantemperate and tropical |
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Discussion | The records for Rhode Island and Wisconsin are according to M. L. Horvat (1941); we have not seen specimens from those states. Cyperus echinatus is usually recognized by its tight, nearly spheric spikes; it may occasionally be hard to distinguish from C. croceus and C. retrorsus. Compared to C. retrorsus, C. echinatus has larger spikelets and longer floral scales, anthers, and achenes. In contrast to C. echinatus, C. croceus has looser spikes, shorter, broader, greenish or yellowish floral scales, shorter, more ovoid achenes, and shorter anthers. Furthermore, C. echinatus is predominantly an inland species of roadsides, pastures, and other disturbed ground; C. retrorsus is primarily a coastal species and occurs in drier, sandier sites. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 600 (96 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 182. | FNA vol. 23, p. 141. | ||||||||||||||||
Parent taxa | Cyperaceae > Cyperus > subg. Cyperus | Cyperaceae | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
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Synonyms | Scirpus echinatus, C. ovularis, C. ovularis var. americanus, C. ovularis var. sphaericus, C. ovularis var. wolfii, C. wolfii, Kyllinga ovularis, Mariscus ovularis | |||||||||||||||||
Name authority | (Linnaeus) Alph. Wood: Class-book Bot. ed. s.n.(b), 734. (1861) | Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754) | ||||||||||||||||
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