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globe flatsedge, teasel sedge

cyperus, flatsedge, galingale, nutsedge, umbrella-sedge

Habit Herbs, perennial, single-stemmed to loosely cespitose. Herbs, perennial or less often annual, cespitose or not, rhizomatous, stoloniferous, rarely tuberous.
Culms

basally cormlike, trigonous, (15–)30–100 cm × 0.5–3.5 mm, glabrous.

solitary or not, trigonous or round, glabrous or scabridulous with extrorse or antrorse (rarely retrorse) prickles.

Leaves

flat to V-shaped, 10–65 cm × 3–9 mm, adaxial surface, margins minutely scabridulous.

usually basal;

ligules absent;

blades keeled abaxially, flat, V-, or inversely W-shaped in cross section.

Inflorescences

spikes densely globose to globose-ovoid, 8–17 mm wide;

rays 3–12, 2–12 cm, scaberous adaxially especially distally;

rachis 4–8 mm;

bracts (3–)4–7, ascending at 30(–45)°, flat, 5–35 cm × 2–9 mm;

rachilla persistent, wings 0.5–0.7 mm wide.

terminal, rarely pseudolateral, 1st order subumbellate to capitate, 2d order with spicate or digitately arranged spikelets, rarely a solitary spikelet;

spikelets 1–150; 1st order rays unequal (rarely equal) in length, produced singly from the axils of inflorescence bracts;

involucral bracts 1–22, spirally arranged at culm apex, spreading to erect, leaflike.

Spikelets

50–100, oblong-lanceoloid, ± terete-quadrangular, (3.5–)4–7 × 1–1.4 mm;

distal spikelet spreading or ascending;

floral scales persistent, 3–5, appressed, stramineous to brownish, 4-ribbed laterally, oblong-elliptic, 3.5–4.5 × 1–1.8 mm, membranous, apex entire or emarginate with mucro to 0.3 mm.

scales to 76, distichous, each subtending flower, cylindric to compressed, borne spicately or digitately at ends of rays (occasionally proliferous).

Flowers

anthers 0.4–0.8 mm;

styles 0.5–0.6 mm;

stigmas 1 mm.

bisexual [rarely unisexual], in axils of distichous floral scales, bases often decurrent onto rachilla as ± hyaline wings;

perianth absent;

stamens 1–3;

styles linear, 2–3-fid, base deciduous or persistent;

stigmas 2–3.

Achenes

brown, ± stipitate, oblong, (1.5–)1.8–2.3 × 0.5–0.6(–0.7) mm (1/2 length of floral scales), apex obtuse, surfaces puncticulate.

biconvex, flattened, or trigonous.

Cyperus echinatus

Cyperus

Phenology Fruiting summer–early fall.
Habitat Disturbed, sunny sites, in mesic places, well-drained soils
Elevation 0–500 m (0–1600 ft)
Distribution
from FNA
AL; AR; DC; DE; FL; GA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; West Indies
[WildflowerSearch map]
[BONAP county map]
from USDA
Pantemperate and tropical
[BONAP county map]
Discussion

The records for Rhode Island and Wisconsin are according to M. L. Horvat (1941); we have not seen specimens from those states.

Cyperus echinatus is usually recognized by its tight, nearly spheric spikes; it may occasionally be hard to distinguish from C. croceus and C. retrorsus. Compared to C. retrorsus, C. echinatus has larger spikelets and longer floral scales, anthers, and achenes. In contrast to C. echinatus, C. croceus has looser spikes, shorter, broader, greenish or yellowish floral scales, shorter, more ovoid achenes, and shorter anthers. Furthermore, C. echinatus is predominantly an inland species of roadsides, pastures, and other disturbed ground; C. retrorsus is primarily a coastal species and occurs in drier, sandier sites.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 600 (96 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas 2 (3 in C. serotinus); achenes biconvex.
→ 2
1. Stigmas 3; achenes trigonous, plano-convex, or terete.
→ 3
2. Achenes laterally flattened, borne with edge toward rachilla.
subg. Pycreus
2. Achenes dorsiventrally flattened, borne with face toward rachilla.
subg. Juncellus
3. Spikelets borne in digitate clusters (rarely singly) or in umbellate or glomerulate heads.
subg. Pycnostachys
3. Spikelets borne in spikes on conspicuous rachis.
→ 4
4. Rachilla continuous or articulate only at base.
subg. Cyperus
4. Rachilla articulate at base of each scale, mature spikelet disarticulating into segments consisting of scale, internode, and rachilla wings.
subg. Diclidium
Source FNA vol. 23, p. 182. FNA vol. 23, p. 141. Authors: Gordon C. Tucker*, Brian G. Marcks*, J. Richard Carter *.
Parent taxa Cyperaceae > Cyperus > subg. Cyperus Cyperaceae
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Subordinate taxa
C. subg. Cyperus, C. subg. Diclidium, C. subg. Juncellus, C. subg. Pycnostachys, C. subg. Pycreus
Synonyms Scirpus echinatus, C. ovularis, C. ovularis var. americanus, C. ovularis var. sphaericus, C. ovularis var. wolfii, C. wolfii, Kyllinga ovularis, Mariscus ovularis
Name authority (Linnaeus) Alph. Wood: Class-book Bot. ed. s.n.(b), 734. (1861) Linnaeus: Sp. Pl. 1: 44. (1753): Gen. Pl. ed. 5, 26. (1754)
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