Cyperus drummondii |
Cyperus sphaerolepis |
|
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Drummond's sedge |
Rusby's flatsedge, Rusby's sedge |
|
Habit | Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. | Herbs, perennial, rhizomatous. |
Culms | trigonous, 35–170 cm × 2–4.7 mm, scabrid on angles. |
basally cormlike, trigonous, (8–)15–40(–60) cm × (0.4–)0.8–1.5(–2) mm, glabrous or sparsely to densely scabridulous to minutely scabrid on angles or usually on surface, just proximal to bracts or over distal few centimeters. |
Leaves | 4–12, with readily visible cross ribs especially on abaxial surface, flat to V-shaped, 25–100 cm × 2.4–11 mm. |
(1–)3–4, V-shaped, (7–)15–25(–40) cm × 1–3(–5) mm, sparsely minutely scabridulous on margins and keel in distal 1/3, otherwise glabrous. |
Inflorescences | heads digitate, globose, 1–1.7 cm diam.; rays 3–5, 1–7 cm; 2d order rays absent; bracts 3–5, horizontal to ascending at 30°, with prominent cross ribs, V-shaped, (4–)11–35(–48) cm × 1–6.7 mm (longest 1/2 as long as culm); 2d order bracts absent. |
spikes 1–3, sessile (occasionally 1 spike on peduncle 5–20 cm, produced from axil of distalmost leaf), loosely cylindric to oblong-ovoid, (9–)15–30(–45) × (3–)8–15(–18) mm; bracts 2–3(–7), longest erect, or very nearly so, others spreading, (1–)3–20(–26) cm × (0.5–)1–3(–5) mm; rays (1–)2–6(–11), (1–)2–7(–12) cm, in some depauperate plants rays less than 1 cm; rachilla ± deciduous, wingless. |
Spikelets | 10–40, oblong to linear-lanceoloid, 5–1.8 × 1.5–2.2 mm; floral scales 18–42, pale grayish green, turning brownish, proximally greenish or brownish, 2-keeled, weakly to distinctly 1-ribbed, proximal 1/2 2-ribbed, ovate, 1.4–1.6 × 0.9–1.4 mm, acute (sometimes mucronulate), apically glabrous, occasionally scabridulous. |
(4–)8–25(–35), oblong-ellipsoid to linear, compressed-quadrangular, (2–)7–11(–15) × (2–)2.4–3.2(–3.5) mm; floral scales (1–)4–8(–14), laterally dull whitish, stramineous, or light brown, or reddish brown to deep red, often red-dotted especially near keel, medially greenish, broadly ovate-deltate to ± orbiculate, laterally strongly 3(–4)-ribbed, medially weakly 1–3-ribbed, (1.6–)1.8–2.4 × (1.8–)2.2–2.8 mm, margins clear, erose especially apically, apex obtuse, tipped with straight mucro 0.1–0.2(–0.3) mm. |
Flowers | stamens 1–2; anthers 0.8–1.2 mm; styles 0.8–1 mm; stigmas 0.6–0.8 mm. |
anthers 0.4–0.6 mm; styles 0.4(–0.8) mm; stigmas 0.6–1 mm. |
Achenes | brown, slightly stipitate, oblong-ellipsoid (2.4–3.9 times longer than wide), 1–1.2 × 0.3–0.5 mm, apical beak 0.1–0.5 mm, surfaces glabrous. |
finely reticulate with isodiametric cells, this layer gradually wearing away to reveal inner brown to reddish brown layer, stipitate, ovoid to obovoid, 1.4–1.6(–1.9) × 1–1.3(–1.4) mm, base cuneate to ± attenuate, stipe 0.1–0.2 × 0.2 mm, apex obtuse, slightly apiculate, inner layer papillose. |
Cyperus drummondii |
Cyperus sphaerolepis |
|
Phenology | Fruiting late spring–summer. | Fruiting summer. |
Habitat | Wet, relatively undisturbed habitats, especially flatwoods, ponds, seepage slopes, coastal prairies | Clearings in montane forests |
Elevation | 0–200 m (0–700 ft) | 1000–2500 m (3300–8200 ft) |
Distribution |
AL; FL; GA; LA; MS; SC; TX; Central America; South America; West Indies |
AZ; NM; TX; Mexico |
Discussion | Cyperus drummondii has been treated as a synonym (G. C. Tucker 1994), or a variety (G. Kükenthal 1935–1936), or a subspecies of C. virens (M. F. Denton 1978b). Recent quantitative and field studies (J. R. Carter et al. 1999) provided compelling evidence that specific status is appropriate. Compared to Cyperus virens, C. drummondii is a taller species of less disturbed habitats and has narrower spikelets, ovate scales, and fewer inflorescence bracts. Cyperus virens and C. drummondii are readily distinguished from other Cyperus species in the United States by their sharply angled, scabrid culms and conspicuously septate leaf blades. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus sphaerolepis has long been known by the synonym C. rusbyi and has been treated as a variety of C. fendlerianus. Specific status for C. sphaerolepis is supported by B. G. Marcks (1972) and G. C. Tucker (1994). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 177. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. robustus, C. virens subsp. drummondii, C. virens var. drummondii, C. virens var. robustus | C. fendlerianus var. debilis, C. fendlerianus var. leucolepis, C. leucolepis, C. rusbyi, C. schweinitzii var. debilis |
Name authority | Torrey & Hooker: Ann. Lyceum Nat. Hist. New York 3: 437, errata. (1836) | Boeckeler: Linnaea 35: 609. (1868) |
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