Cyperus drummondii |
Cyperus granitophilus |
|
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Drummond's sedge |
granite flatsedge |
|
Habit | Herbs, perennial, cespitose; rhizomes 0.5–2 cm, often absent. | Herbs, annual, cespitose, with roots fibrous. |
Culms | trigonous, 35–170 cm × 2–4.7 mm, scabrid on angles. |
trigonous, glabrous. |
Leaves | 4–12, with readily visible cross ribs especially on abaxial surface, flat to V-shaped, 25–100 cm × 2.4–11 mm. |
V-shaped to flat, (2–)4–11 cm × 3–4 mm. |
Inflorescences | heads digitate, globose, 1–1.7 cm diam.; rays 3–5, 1–7 cm; 2d order rays absent; bracts 3–5, horizontal to ascending at 30°, with prominent cross ribs, V-shaped, (4–)11–35(–48) cm × 1–6.7 mm (longest 1/2 as long as culm); 2d order bracts absent. |
rays absent; bracts 2–4, ± horizontal, flat, 1–6.5 cm × 2–3 mm; rachilla persistent, wingless. |
Spikelets | 10–40, oblong to linear-lanceoloid, 5–1.8 × 1.5–2.2 mm; floral scales 18–42, pale grayish green, turning brownish, proximally greenish or brownish, 2-keeled, weakly to distinctly 1-ribbed, proximal 1/2 2-ribbed, ovate, 1.4–1.6 × 0.9–1.4 mm, acute (sometimes mucronulate), apically glabrous, occasionally scabridulous. |
10–40, compressed, ovoid to linear-lanceoloid, quadrangular, 5–9 × 2–3 mm; floral scales deciduous, 5–20, spreading, pale greenish to reddish brown, laterally (5–)7–9(–13)-ribbed, ovate, 3–4 × 1–2 mm, apex straight, excurved mucronate, or cuspidate. |
Flowers | stamens 1–2; anthers 0.8–1.2 mm; styles 0.8–1 mm; stigmas 0.6–0.8 mm. |
stamens 1–2; filaments 1.5 mm; anthers 0.8–1 mm, connective apex reddish yellow, less than 0.1 mm; styles 0.8–1 mm; stigmas 1–1.2 mm. |
Achenes | brown, slightly stipitate, oblong-ellipsoid (2.4–3.9 times longer than wide), 1–1.2 × 0.3–0.5 mm, apical beak 0.1–0.5 mm, surfaces glabrous. |
brown to reddish brown, broadly ellipsoid to obovoid, 0.8–1.2 × 0.5–0.6 mm, base ± cuneate, apex truncate-obtuse, surfaces puncticulate. |
Cyperus drummondii |
Cyperus granitophilus |
|
Phenology | Fruiting late spring–summer. | Fruiting summer. |
Habitat | Wet, relatively undisturbed habitats, especially flatwoods, ponds, seepage slopes, coastal prairies | Thin soil over outcrops of granite or sandstone of the Piedmont and inner coastal plain |
Elevation | 0–200 m (0–700 ft) | 50–400 m (200–1300 ft) |
Distribution |
AL; FL; GA; LA; MS; SC; TX; Central America; South America; West Indies |
AL; GA; NC; SC; TN; VA
|
Discussion | Cyperus drummondii has been treated as a synonym (G. C. Tucker 1994), or a variety (G. Kükenthal 1935–1936), or a subspecies of C. virens (M. F. Denton 1978b). Recent quantitative and field studies (J. R. Carter et al. 1999) provided compelling evidence that specific status is appropriate. Compared to Cyperus virens, C. drummondii is a taller species of less disturbed habitats and has narrower spikelets, ovate scales, and fewer inflorescence bracts. Cyperus virens and C. drummondii are readily distinguished from other Cyperus species in the United States by their sharply angled, scabrid culms and conspicuously septate leaf blades. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus granitophilus is documented as an autotetraploid derivative of C. squarrosus (L. W. Garoni and W. H. Murdy 1964). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 165. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. robustus, C. virens subsp. drummondii, C. virens var. drummondii, C. virens var. robustus | |
Name authority | Torrey & Hooker: Ann. Lyceum Nat. Hist. New York 3: 437, errata. (1836) | McVaugh: Castanea 2: 103, figs. 4–8. (1937) |
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