Cyperus difformis |
Cyperus rotundus |
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Asian flatsedge, smallflower umbrella sedge, variable flatsedge |
chaguan humatag, coco-grass, nutgrass, pakopako, purple nut-sedge |
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Habit | Herbs, annual, cespitose. | Herbs, perennial, stoloniferous; stolons (2–)5–12 cm × 1–2 mm, bearing tubers 3–8(–12) mm diam., wiry, springy when dried, indurate. |
Culms | 1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous. |
trigonous, 10–35(–40) cm × 0.7–3.4 mm, basally indurate, glabrous. |
Leaves | 2–7, flat, (2–)7–22 cm × 2.2–4 mm. |
V-shaped to flanged V-shaped, 5–30 cm × 2–6 mm. |
Inflorescences | heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.; rays 1–5, 2–32 mm; bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous. |
spikes 1(–3), broadly ellipsoid, (12–)15–25(–30) × (12–)20–30(–50) mm, rays (3–)4–6(–7), 0.2–10 cm;, bracts (2–)3–5, horizontal to ascending at 45°, V-shaped to flanged V-shaped, 0.5–10 cm × 0.5–4 mm, rachilla persistent, wings 0.5–1 mm wide. |
Spikelets | 30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm; floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate. |
(2–)3–7(–12), compressed, linear, 4–40 × 1.3–1.8 mm, floral scales persistent, 6–36(–42), spreading or appressed, purple to reddish brown, with narrow clear border and green midrib, 7–9-ribbed, ovate, (1.8–)2.6–3.4 × 2.2–3 mm, apex obtuse. |
Flowers | stamens 1 or 2; anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged; styles 0.1 mm; stigmas 0.1–0.3 mm. |
anthers 1–2.5 mm, styles 1.3–3.5 mm, stigmas (1.8–)2–3.3 mm. |
Achenes | light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose. |
black, sessile, ellipsoid, abaxial face convex, adaxial face concave, 1.4–1.7(–1.9) × 0.8–1 mm, apex obtuse, surfaces puncticulate. |
Cyperus difformis |
Cyperus rotundus |
|
Phenology | Fruiting summer. | Fruiting summer–fall. |
Habitat | Disturbed, muddy soils, shallow waters | Croplands, disturbed soils usually |
Elevation | 0–1000 m (0–3300 ft) | 0–400 m (0–1300 ft) |
Distribution |
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
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AL; AR; AZ; CA; FL; GA; LA; MO; MS; NC; NM; SC; TN; TX; VA; Mexico; Central America; South America; Asia; Africa; Australia
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Discussion | Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus rotundus is documented in Connecticut, Delaware, New York, and Pennsylvania; there is no evidence of persistent populations. Cyperus esculentus and C. rotundus are the only two species of subg. Cyperus in the New World that produce tuberiferous stolons. The two species also have persistent floral scales and persistent rachillas, a combination of characteristics not found in any other New World species of Cyperus. Cyperus rotundus is distinguished from other species of the genus in the New World by its open spikes composed of linear reddish spikelets borne on a conspicuous slender rachis. Cyperus rotundus is usually acknowledged to be the world’s worst weed (cf. G. C. Tucker 1987). In the United States, it does not grow north of the mean 1°C January isotherm. Cyperus esculentus (preceding species) is a serious weed in much of the world, especially in cooler regions where the more tropical C. rotundus does not grow. Cyperus esculentus is able to tolerate lower air temperatures (as low as -18°C). The two species apparently differ also in their thermal optima for growth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 156. | FNA vol. 23, p. 169. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. lateriflorus | |
Name authority | Linnaeus: Cent. Pl. II, 6. (1756) | Linnaeus: Sp. Pl. 1: 45. (1753) |
Web links |