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Asian flatsedge, smallflower umbrella sedge, variable flatsedge

Egyptian paper-reed, papyrus

Habit Herbs, annual, cespitose. Herbs, perennial, rhizomatous, stout.
Culms

1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous.

roundly trigonous, 300–500 cm × 15–45 mm, glabrous.

Leaves

2–7, flat, (2–)7–22 cm × 2.2–4 mm.

bladeless.

Inflorescences

heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.;

rays 1–5, 2–32 mm;

bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous.

spikes loosely cylindric, 10–20 × 6–10 mm;

rays 40–100, drooping or arching, slender, 10–30 cm; 2d order rays 8–20 cm;

bracts 4–10, ± erect, V-shaped, 3–8 cm × 4–15 mm; 2d order bracts 2–5, (1.5–)4–16 cm × 0.5–2 mm;

rachilla persistent, separating laterally, remaining firmly attached basally, wings 0.3–0.4 mm wide.

Spikelets

30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm;

floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate.

6–30, slightly compressed, linear, ± quadrangular, 6–10 × 0.8–1 mm;

floral scales 6–16, reddish beside 5-ribbed green medial part, white to hyaline near margins, ovate-elliptic, 1.8–2.2 × 1.2–1.5 mm, apex acute to obtuse.

Flowers

stamens 1 or 2;

anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged;

styles 0.1 mm;

stigmas 0.1–0.3 mm.

anthers 0.8–1 mm (connectives prolonged beyond anther as red subulate appendage 0.2–0.5 mm, apex usually setose);

styles 0.2–0.4 mm;

stigmas 0.8–1.2 mm.

Achenes

light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose.

pale brown, sessile, oblong, 0.8–1 × 0.4 mm, apex scarcely apiculate, surfaces puncticulate.

Cyperus difformis

Cyperus papyrus

Phenology Fruiting summer. Fruiting summer.
Habitat Disturbed, muddy soils, shallow waters Stream banks, marshes
Elevation 0–1000 m (0–3300 ft) 0–30 m (0–100 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; s Europe; sw Asia; Africa [Introduced in North America]
[BONAP county map]
Discussion

Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyperus papyrus is conspicuous in the field by its great height, leafless culm, and open, hemispheric inflorescence with drooping rays.

Ancient Egyptians, Greeks, and Romans prepared paper from the pith of the culms of Cyperus papyrus, which is abundant along the Nile River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 156. FNA vol. 23, p. 173.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Synonyms C. lateriflorus
Name authority Linnaeus: Cent. Pl. II, 6. (1756) Linnaeus: Sp. Pl. 1: 47. (1753)
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