Cyperus difformis |
Cyperus lupulinus |
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Asian flatsedge, smallflower umbrella sedge, variable flatsedge |
Great Plains flatsedge, slender sand sedge |
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Habit | Herbs, annual, cespitose. | Herbs, perennial, cespitose, bases cormose; rhizomes knotted, beaded. | ||||
Culms | 1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous. |
trigonous, (3–)10–50 cm × 0.4–1.2 mm, glabrous. |
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Leaves | 2–7, flat, (2–)7–22 cm × 2.2–4 mm. |
flat, 5–40 cm × 1–3.5 mm. |
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Inflorescences | heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.; rays 1–5, 2–32 mm; bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous. |
spikes rather densely ovoid to globose, 1.2–3.5 cm; rays 0 or 1–4, 1–6 cm; rachis 1–3.5 mm; bracts 2–4, horizontal to reflexed, flat, 6–25 cm; rachilla ± deciduous, wingless. |
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Spikelets | 30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm; floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate. |
15–60, compressed, oblong-lanceoloid, (3–)6–22 × 2.5–4 mm; floral scales deciduous, 5–22, off-white to light reddish brown, laterally 3–5-ribbed, ovate-elliptic, 2.5–4 × 2–2.6 mm, margins loosely spreading or clasping achene, apex entire to mucronate, mucro 0.05–0.2 mm. |
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Flowers | stamens 1 or 2; anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged; styles 0.1 mm; stigmas 0.1–0.3 mm. |
anthers 0.3–0.6 mm; styles 1 mm; stigmas 1–1.5 mm. |
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Achenes | light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose. |
dark brown or black, sessile, oblong-ellipsoid to ellipsoid, 1.7–2.2 × 0.8–1.2 mm, apex obtuse, apiculate, surfaces puncticulate. |
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2n | = 166. |
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Cyperus difformis |
Cyperus lupulinus |
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Phenology | Fruiting summer. | |||||
Habitat | Disturbed, muddy soils, shallow waters | |||||
Elevation | 0–1000 m (0–3300 ft) | |||||
Distribution |
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
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AR; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OR; PA; RI; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
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Discussion | Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Cyperus lupulinus was studied in detail (B. G. Marcks 1974). Ranges of the two subspecies overlap somewhat. Cyperus lupulinus subsp. lupulinus is found chiefly in the Great Plains, and subsp. macilentus is centered in the Northeast. It is seldom difficult to assign specimens to subspecies. The hybrid of Cyperus lupulinus with C. schweinitzii is C. ×mesochorus Geise. It is occasionally encountered with the two parent species in the north-central states (Illinois, Indiana, Michigan, Minnesota, and Wisconsin) and has been recorded once from Quebec. The hybrid is similar in size to C. schweinitzii; it has fewer rays, inflorescence bracts 30–45º above horizontal, and floral scales with mucros 0.4–0.5 mm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 23, p. 156. | FNA vol. 23, p. 176. | ||||
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | C. lateriflorus | Scirpus lupulinus | ||||
Name authority | Linnaeus: Cent. Pl. II, 6. (1756) | (Sprengel) Marcks: Trans. Wisconsin Acad. Sci. 62: 271. (1974) | ||||
Web links |
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