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Asian flatsedge, smallflower umbrella sedge, variable flatsedge

Great Plains flatsedge, slender sand sedge

Habit Herbs, annual, cespitose. Herbs, perennial, cespitose, bases cormose; rhizomes knotted, beaded.
Culms

1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous.

trigonous, (3–)10–50 cm × 0.4–1.2 mm, glabrous.

Leaves

2–7, flat, (2–)7–22 cm × 2.2–4 mm.

flat, 5–40 cm × 1–3.5 mm.

Inflorescences

heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.;

rays 1–5, 2–32 mm;

bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous.

spikes rather densely ovoid to globose, 1.2–3.5 cm;

rays 0 or 1–4, 1–6 cm;

rachis 1–3.5 mm;

bracts 2–4, horizontal to reflexed, flat, 6–25 cm;

rachilla ± deciduous, wingless.

Spikelets

30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm;

floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate.

15–60, compressed, oblong-lanceoloid, (3–)6–22 × 2.5–4 mm;

floral scales deciduous, 5–22, off-white to light reddish brown, laterally 3–5-ribbed, ovate-elliptic, 2.5–4 × 2–2.6 mm, margins loosely spreading or clasping achene, apex entire to mucronate, mucro 0.05–0.2 mm.

Flowers

stamens 1 or 2;

anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged;

styles 0.1 mm;

stigmas 0.1–0.3 mm.

anthers 0.3–0.6 mm;

styles 1 mm;

stigmas 1–1.5 mm.

Achenes

light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose.

dark brown or black, sessile, oblong-ellipsoid to ellipsoid, 1.7–2.2 × 0.8–1.2 mm, apex obtuse, apiculate, surfaces puncticulate.

2n

= 166.

Cyperus difformis

Cyperus lupulinus

Phenology Fruiting summer.
Habitat Disturbed, muddy soils, shallow waters
Elevation 0–1000 m (0–3300 ft)
Distribution
from FNA
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; OR; PA; RI; TN; TX; VA; VT; WI; WV; NB; NS; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Cyperus lupulinus was studied in detail (B. G. Marcks 1974). Ranges of the two subspecies overlap somewhat. Cyperus lupulinus subsp. lupulinus is found chiefly in the Great Plains, and subsp. macilentus is centered in the Northeast. It is seldom difficult to assign specimens to subspecies.

The hybrid of Cyperus lupulinus with C. schweinitzii is C. ×mesochorus Geise. It is occasionally encountered with the two parent species in the north-central states (Illinois, Indiana, Michigan, Minnesota, and Wisconsin) and has been recorded once from Quebec. The hybrid is similar in size to C. schweinitzii; it has fewer rays, inflorescence bracts 30–45º above horizontal, and floral scales with mucros 0.4–0.5 mm.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Floral scales 2.5–4 mm, margins not clasping achene; spikelets with 5–22 floral scales.
subsp. lupulinus
1. Floral scales 1.8–2.5 mm, margins tightly clasping achene; spikelets with 3–7 floral scales.
subsp. macilentus
Source FNA vol. 23, p. 156. FNA vol. 23, p. 176.
Parent taxa Cyperaceae > Cyperus > subg. Pycnostachys Cyperaceae > Cyperus > subg. Cyperus
Sibling taxa
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. lupulinus, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
C. acuminatus, C. aggregatus, C. alopecuroides, C. amabilis, C. articulatus, C. bipartitus, C. cephalanthus, C. compressus, C. croceus, C. cuspidatus, C. dentatus, C. dentoniae, C. diandrus, C. difformis, C. digitatus, C. dipsaceus, C. distans, C. distinctus, C. drummondii, C. echinatus, C. elegans, C. entrerianus, C. eragrostis, C. erythrorhizos, C. esculentus, C. fendlerianus, C. filicinus, C. filiculmis, C. filiformis, C. flavescens, C. flavicomus, C. floribundus, C. fugax, C. fuligineus, C. fuscus, C. giganteus, C. gracilis, C. granitophilus, C. grayi, C. grayoides, C. haspan, C. hermaphroditus, C. houghtonii, C. hypopitys, C. hystricinus, C. involucratus, C. iria, C. laevigatus, C. lancastriensis, C. lanceolatus, C. lecontei, C. lentiginosus, C. ligularis, C. manimae, C. microiria, C. mutisii, C. niger, C. ochraceus, C. odoratus, C. onerosus, C. ovatus, C. owanii, C. oxylepis, C. pallidicolor, C. papyrus, C. parishii, C. pilosus, C. planifolius, C. plukenetii, C. polystachyos, C. prolifer, C. prolixus, C. pseudothyrsiflorus, C. pseudovegetus, C. pumilus, C. reflexus, C. refractus, C. retroflexus, C. retrofractus, C. retrorsus, C. rotundus, C. sanguinolentus, C. schweinitzii, C. serotinus, C. seslerioides, C. setigerus, C. spectabilis, C. sphacelatus, C. sphaerolepis, C. squarrosus, C. strigosus, C. surinamensis, C. tetragonus, C. thyrsiflorus, C. virens
Subordinate taxa
C. lupulinus subsp. lupulinus, C. lupulinus subsp. macilentus
Synonyms C. lateriflorus Scirpus lupulinus
Name authority Linnaeus: Cent. Pl. II, 6. (1756) (Sprengel) Marcks: Trans. Wisconsin Acad. Sci. 62: 271. (1974)
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