Cyperus difformis |
Cyperus hystricinus |
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Asian flatsedge, smallflower umbrella sedge, variable flatsedge |
bristly flatsedge |
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Habit | Herbs, annual, cespitose. | Herbs, perennial, cespitose, with well-developed rhizomes. |
Culms | 1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous. |
trigonous, 30–100 cm × 2–4 mm, basally cormlike, glabrous. |
Leaves | 2–7, flat, (2–)7–22 cm × 2.2–4 mm. |
flat to broadly V-shaped, 20–70 cm × 4–6 mm, glabrous except for marginal prickles. |
Inflorescences | heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.; rays 1–5, 2–32 mm; bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous. |
spikes loosely to densely ovoid, oblong (2 times long as wide), 10–12 × 6–9 mm; rays 6–11, 1–16 cm, glabrous; bracts 5–10, ascending at 30–45°, flat, 6–25 cm × 3–6 mm; rachilla persistent, wings 0.5 mm wide, covering nearly entire length of achene. |
Spikelets | 30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm; floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate. |
(20–)40–100(–120), proximal spikelets reflexed somewhat, distal ones divaricate, ± terete, lanceoloid, 6–6.8 × 1–1.4 mm; floral scales persistent, 1–2(–3), appressed, golden brown, lanceolate, laterally 5–6-ribbed, 3.8–4.9 × 1.4–1.6 mm, apex mucronate, mucro at most 0.3 mm. |
Flowers | stamens 1 or 2; anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged; styles 0.1 mm; stigmas 0.1–0.3 mm. |
anthers 1–1.3 mm; styles 0.8–1 mm; stigmas 2–3 mm. |
Achenes | light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose. |
brown, sessile, linear, 2.5–2.8 × 0.5–0.7 mm, apex obtuse, apiculate, surfaces muriculate. |
Cyperus difformis |
Cyperus hystricinus |
|
Phenology | Fruiting summer. | Fruiting late summer–early fall (Jul–Sep). |
Habitat | Disturbed, muddy soils, shallow waters | Xeric, sandy soils of sand hills and pine barrens |
Elevation | 0–1000 m (0–3300 ft) | 0–200 m (0–700 ft) |
Distribution |
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
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AL; AR; DC; DE; GA; LA; MD; MO; MS; NC; NJ; TX; VA
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Discussion | Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyperus hystricinus resembles C. plukenetii and C. retrofractus; it can be readily distinguished from both by its glabrous culms. Overly mature specimens of C. lancastriensis are frequently misidentified as C. hystricinus; C. hystricinus may be confirmed by its narrow, nearly glabrous leaves and bracts, golden brown spikelets, longer, narrower achenes, and elongated rhizome internodes (to 15 mm vs. 5 mm in 76. C. lancastriensis). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 156. | FNA vol. 23. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. lateriflorus | C. retrofractus var. hystricinus |
Name authority | Linnaeus: Cent. Pl. II, 6. (1756) | Fernald: Rhodora 8: 127. (1906) |
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