Cyperus difformis |
Cyperus fendlerianus |
|
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Asian flatsedge, smallflower umbrella sedge, variable flatsedge |
Fendler flatsedge, Fendler's flatsedge, Fendler's nutgrass, Fendler's sedge |
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Habit | Herbs, annual, cespitose. | Herbs, perennial, rhizomatous. |
Culms | 1–15, trigonous, 7–30 cm × 1.2–2.5 mm, soft (flattened in pressing), glabrous. |
trigonous, (7–)20–70(–85) cm × (0.5–)1–1.8(–2.6) mm, glabrous or sparsely to densely scabridulous on surfaces or on angles immediately proximal to inflorescence in distal 3–6 cm. |
Leaves | 2–7, flat, (2–)7–22 cm × 2.2–4 mm. |
2–3, flat to V-shaped, (12–)20–40(–55) cm × (2–)3–5(–7) mm. |
Inflorescences | heads dense, 7–17 mm diam.; when rays short, heads sessile or nearly so, then densely irregularly lobate, 12–35 mm diam.; rays 1–5, 2–32 mm; bracts 2–4, longest bract erect or nearly so, appearing as continuation of culm, other bracts horizontal to ascending, 1–22 cm × 0.5–3.5 mm, margins and keel minutely scabridulous. |
spikes (1–)3–4(–6), largest erect, others at base of central spike, erect spike sessile, loosely to densely cylindric to ovoid-cylindric, (12–)18–30 × (6–)12–20 mm, remaining spikes 1/2 as large; rays absent; bracts (2–)3–6(–10), ± horizontal to reflexed parallel to culm, (2–)6–16(–25) cm × (1.5–)2–4(–7) mm; rachilla ± deciduous, wingless. |
Spikelets | 30–120, greenish brown to purplish brown, oblong-ellipsoid, compressed, (2–)3–5(–6) × 0.8–1.2 mm; floral scales (6–)12–20(–30), laterally clear margins, stramineous to deep purple, medially greenish, stramineous, or purplish, laterally ribless, medially 3-ribbed, obovate to orbiculate, 0.6–0.8 × 0.6–0.8 mm, apex mucronulate. |
(8–)15–30, spreading to ascending-appressed, oblong-lanceoloid, quadrangular, slightly compressed, 5–8(–10) × (2–)2.4–3.4(–4.2) mm; floral scales (2–)4–6(–8), yellowish, reddish, or greenish brown, medially greenish, straight to excurved, laterally (2–)3–4(–5)-ribbed, medially (1–)3–(–5)-ribbed, ovate-orbiculate, 2.4–2.8(–3.4) × 2.4–2.8(–3.4) mm, apex obtuse, mucronate to cuspidate, stout, (0.1–)0.3–0.6 mm, medially glabrous. |
Flowers | stamens 1 or 2; anthers ovoid-ellipsoid, 0.1 mm, connective not prolonged; styles 0.1 mm; stigmas 0.1–0.3 mm. |
anthers 0.4–0.8 mm; styles 0.7–1.3 mm; stigmas 0.7–1.2 mm. |
Achenes | light brown, obovoid-ellipsoid, 0.6–0.8 × 0.3–0.4 mm (as long as subtending scale), base cuneate, apex obtuse, apiculate, surfaces finely reticulate, papillose. |
finely reticulate, brown to reddish brown, ± stipelike, obovoid, 1.6–1.9 × 1.1–1.4(–1.6) mm, base attenuate to cuneate, 0.1–0.2 × 0.2 mm, apex obtuse, apiculate, surfaces glabrous. |
Cyperus difformis |
Cyperus fendlerianus |
|
Phenology | Fruiting summer. | Fruiting summer. |
Habitat | Disturbed, muddy soils, shallow waters | Clearings in montane forests |
Elevation | 0–1000 m (0–3300 ft) | 500–1800 m (1600–5900 ft) |
Distribution |
AL; AZ; CA; FL; GA; ID; KY; LA; MS; NC; NJ; NM; OR; PA; SC; TN; TX; VA; Mexico; South America; West Indies (Puerto Rico); Central America (Nicaragua, Panama); Eurasia; Indian Ocean Islands; Pacific Islands; Australia [Introduced in North America]
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AZ; CO; NM; TX; WY; Mexico
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Discussion | Cyperus difformis is naturalized in the New World and native to the Old World, where it ranges from southern Europe to southern Africa and eastward to Southeast Asia and Australia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Hybridization between Cyperus fendlerianus and C. sphaerolepis occurs occasionally in the southwestern United States and northern Mexico (B. G. Marcks 1972). The unnamed hybrid appears to be fully fertile and can be recognized by morphologic intermediacy. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 156. | FNA vol. 23, p. 177. |
Parent taxa | Cyperaceae > Cyperus > subg. Pycnostachys | Cyperaceae > Cyperus > subg. Cyperus |
Sibling taxa | ||
Synonyms | C. lateriflorus | Chlorocyperus fendlerianus |
Name authority | Linnaeus: Cent. Pl. II, 6. (1756) | Boeckeler: Linnaea 35: 520. (1868) |
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